VI. CATALOGUE OF GENE SYMBOLS FOR WHEAT: 2000 Supplement

R.A. McIntosh 1 , G.E. Hart 2 , K.M. Devos 3 and W.J. Rogers 4.

1 Plant Breeding Institute, The University of Sydney, 107 Cobbitty Rd., Cobbitty, N.S.W., Australia, 2570.
2 Department of Soil & Crop Sciences, Texas A&M University, College Station, Texas, USA, 77843.
3 John Innes Centre, Norwich Research Park, Colney, Norwich, Norfolk, NR4 7UH, UK.
4 Catedra de Genetica y Fitotecnia, Universidad Nacional del Centro de la Provincia de Buenos Aires, 7300 Azul,
Argentina.

The most recent edition of the Catalog appeared in the Proceedings of the 9th International Wheat Genetics Symposium Vol. 5 (Slinkard AE ed, University Extension Press, University of Saskatchewan, Saskatoon, Canada). A modified version is displayed on the Graingenes Website grains@greengenes.cit.cornell.edu.

The 1999 Supplement is included in 1999 Annual Wheat Newsletter, Wheat Information Service, and is listed in the Graingenes Website. The present Supplement will be offered to editors/curators for similar listing.


2000 Supplement

Revisions.


10. Laboratory Designators for DNA markers

 cdc
 Gusta, L.V.
Crop Development Centre*
University of Saskatchewan
51 Campus Drive
Saskatoon, Saskatchewan, S7N 5A8
Canada
 ocs
 Kato, K.
Dept. of Crop Science
Obihiro* University of Agriculture and Science*Veterinary Medicine
Obihiro 080-8555
Japan
 csl
 Lagudah, E.S.*
Division of Plant Industry
CSIRO*, GPO Box 1600
Canberra ACT 2601
Australia
 pgh
 Shimosaka, E.
Laboratory of Plant Genecology*
Hokkaido* National Agricultural
Experiment Station
1, Hitsujigaoka, Toyohira-ku
Sapporo, 062-8555
Japan
 kvl  Collinge, D.B.
Section for Plant Pathology Dept. of Plant Biology
Royal Veterinary and Agricultural University*
Thorvaldsenvej 40
1871 Frederiksberg C
Denmark
 rgs
 (Rice etiolated shoot* clones)
Sasaki, T. (see rgc)
 msu
 Talbert, L.E.
Plant Sciences Department
Montana State University Bozeman, MT 59717
USA
 sun
 Sharp, PJ.
Plant Breeding Institute
University of Sydney
PMB 11, Camden NSW 2570
Australia


Morphological, Physiological, Molecular and DNA Traits

Gross Morphology: Spike characteristics

Following the first paragraph insert: More than 20 NILs involving genes affecting a range of traits are described in {0066}. These are not yet incorporated into the Catalogue.

Gross Morphology : Spike Characteristics
3. Sphaerococcum

Revision:
The naturally-occurring sphaerococcum gene in chromosome 3D and various mutant alleles conferring a similar phenotype form a homoeologous series. The sphaerococcoid alleles are either recessive or incompletely dominant. All three mapped loci are closely linked to the respective centromeres {0030}. The "a" alleles are allocated to Chinese Spring or "normal" wheats.

S-A1 {0029}. 3A {0056}. v: CS {0029}.
S-A1a {0029}. v: CS {0029}; common wheats {0029}.
S-A1b {0029}. S3 {0056}. v: MS 1453 {0056}.
ma: Xgwm2-3A(S) - 5.1 cM - S-A1 - 6.6 cM - Xgwm720-3A(L) {0030}.
S-B1 {0029}. 3B {0030}. v: CS {0029}.
S-B1a {0029}. v: CS {0029}; common wheats {0029}.
S-B1b {0029}. S2 {0030}. v: MSK 2452 {0056}; MSK 2454 {0056}.
ma: Xgwm685-3B(S) - 4.2 cM - S-B1 - 0.5 cM - Xgwm566/Xgwm845/cent {0030}.
S-D1 {0029}. 3D {1292,0030}; 3DS {1193,1194}; 3DL {692}. v: CS {0029}.
S-D1a {0029}. v: CS {0029}; common wheats {0029}.
S-D1b {0029}. s1 [sp1 {1286}]. i: S-615*11/T. sphaerococcum var. rotundatum {1500}.
s: CS*7/T. sphaerococcum rubiginosum 3D {1304}.
v: Sphaerococcum wheats {0029}.
S-D1c {0029}. S1 {0056}. v: MS 3287 {0056}.
ma: Xgdm72-3D(S) - 8.0 cM - S-D1 - 2.9 cM - Xgwm456-3D/cent {0030}.

Temporary designation
s2 [sp2 {1286}]. Partially dominant {1286} Sphaerococcum simulator {1286}.
Sphaerococcum-like tetraploid wheats were reported {122,475,1282,1286}, but comparisons between them, or with s2, were not made. Whereas Schmidt & Johnson {1281} reported a single recessive factor controlling the sphaerococcum character in tetraploid wheat, Joppa {621} using the same stock found that two recessive genes were necessary to produce this phenotype.

Ear length (new section under 'Spike characteristics')
QEl.ocs-5A.1 {0068}. 5AL {0068}. v: CS(T. spelta 5A)/CS(Cappelle-Desprez 5A) RI mapping population {0079}.
ma: Associated with Xbcd9 {0068}.

 

Awnedness
1. Dominant Inhibitors
1.1 Hooded
Hd. Add: 'ma: Xcdo1387-4A - 8.2 cM - Hd - 7.2 cM - Xpsr163 {0047}.'

Boron Tolerance
Bo3
. 4A {0012}.

Crossability with Rye, Hordeum and Aegilops spp.
1. Common wheat
Current section
2. Tetraploid wheat
The Chinese tetraploid, Ailanmai, possesses recessive crossability genes on chromosomes 1A, 6A, and 7A with the 6A gene being the least effective {0017}.



DNA Markers

Group 1S

Amendments:

Add:

 Xcnl5-1A [{0059}].    AG10F/AG10R.  
 Xglk301-1B [{0031}].  [Xglk301a-1B {0031}].  pTag301.  (2D, 3D, 5D, 7A,B).
 Xglk317-1B [{0031}].  [Xglk317a-1B {0031}].  pTag317.  (3D, 4D, 5A, 6A).
 Xgwm264-1B.1 [{9929}1,{0003}^2^].  [Xgwm264-1B {9929}, Xgwm264c-1B {0003}].  WMS F264/ WMS R264.  (1B, 3B).
 Xgwm498-1A {0035}^2^.    WMS F498/WMS R498.  (1BL).
 Xgwm656-1A {0035}^2^.    WMS F656/WMS R656.  
 Xkvl901(Chs)-1B,D [{0091}].  [Chs-1B,D {0091}].  pBH72-O8 {0098}.  
 Xkvl902(Fmt)-1B,D [{0091].  [Fmt-1B,D {0091}].  pBH72-F1 {0099}.  
 Xpsr593-1B [{0031}].  [Xpsr593a-1B {0031}].  PSR593.  (2B, 4B, 7B).
 Xpsr642-1B {0031}.    PSR642.  
 Xpsr960-1B [{0031}].    PSR960.  
 Xsfr3(LRR)-1A,B [{0031}].  [CD9a,b-1A {0031}].  CD9.  
.

Group 1L

Amendments:

Add:

 Xkvl903(Chi2)-1B [{0091}].  [Cht22-1B {0091}].  pBH72-N12 {0092}.  
 Xgwm131-1B.2 [{0003}].  [Xgwm131b-1B {0003}].  WMS F131/WMS R131.  (1B, 3B).
 Xgwm636-1B [{0003}].  [Xgwm636b-1B {0003}].  WMS F636/WMS R636.  (2A).
 Xpsr924-1A,B,D {0043}.    PSR924.  (2B).


Group 1

Amendments:

Add:

Xgwm264-1B.2 [{0003}2].  [Xgwm264a-1B {0003}].  WMS F264/WMS R264  (1B, 3B).
 Xpsr967-1A,B {598}.    PSR967.  (4B, 5A, 6AS, 6BL).
 Xwpg501(Pdi)-1B {0064}.    pTAPDI501.  (4A,B,D).

Group 2S

Amendments:


Add:

 Xcsu182(Sod)-2B [{0091}].  [Sod-2B {0091}].  CSU182.  
 Xglk197-2A {0031}1, 2B {9926}^4^.    pTag197.  (7B).
 Xglk222-2A {0031}, 2D {822}.    pTag222.  (5B,D).
 The arm location of Xglk222-2D was not reported in {822}.
 Xglk302-2B [{822, 0049}].  [Xglk302b-2B {822,0049}].  pTag302.  (4A,D).
 The arm location of Xglk302-2B was not reported in {822}.
 Xglk398-2B.1,B.2 [{822,0049}].  [Xglk398a,b-2B {822,0049}].  pTag398.  (7A,B).
  The arm location of Xglk398-2B.1,B.2 was not reported in {822}.
 Xglk400-2B {822,0049}.    pTag400.  
  The arm location of Xglk400-2B was not reported in {822}.
 Xglk407-2B {822,0049}.    pTag407.  (5A).
 The arm location of Xglk407-2B was not reported in {822}.
 Xglk471-2B [{822,0049}].  [Xglk471b-2B {822,0049}.  pTag471.  
 Xglk546-2B.2 [{822,0049}].  [Xglk546f-2B {822,0049}.  pTag546.  (3B, 5A, 6B, 7A, 7B).
  The arm location of Xglk546-2B.2 was not reported in {822}.
 Xglk661-2B [{822,0049}].  [Xglk661c-2B {822,0049}.  pTag661.  (4A,B,D).
  The arm location of Xglk661-2B was not reported in {822}.
 Xglk687-2B {822,0049}.    pTag687.  
  The arm location of Xglk687-2B was not reported in {822}.
 Xglk703-2B {822,0049}.    pTag703.  
  The arm location of Xglk703-2B was not reported in {822}.
 Xglk2002(Hst2a-1)-2A,B,D [{0049}].  [Hst2a-A1,B1,D1 {0049}].  pwcH2A-4.  
 Xgwm71-2B [{0035}]^2^.  [Xgwm71a-2B {0035}.]  WMS F71/WMS R71.  (2AS, 2A, 3D).
 Xgwm122-2A {9929}^1^, {0035}^2^.    WMS F122/WMS R122.  (3B, 5A,B,D,
6AL, 6BS, 7D).
  The arm location of Xgwm122-2A was not reported in {9929}.
 Xgwm275-2A {9929}^1^,{0035}^2^.    WMS F275/WMS R275.  
  The arm location of Xgwm275-2A was not reported in {9929}.
 Xgwm448-2A {9929}^1^,{0035}^2^.    WMS F448/WMS R448.  
 The arm location of Xgwm448-2A was not reported in {9929}.
 Xgwm547-2B {0035}2.    WMS F547/WMS R547.  (3B).
 Xksu904(Per2)-2A,B [{0091}].  [Per2-2A,B {0091}].  pox22.3 {0093}.  
 Xpsr172(Lhcb1)-2B [{0031}].  [Xpsr172-2B {0031}.  PSR172.  (1A, 5A,B,D, 7A,B,D).
 Xpsr386-2A [{0031}].  [Xpsr386c-2A {0031}].  PSR386.  (1A, 3B, 5A, 7A).


Group 2L

Amendments:

Add:

 Xbcd377-2A {0078}.    BCD377.  
 Xcdc2(Sod3)-2A,B,D [{0054}].    SOD3.1 & SOD3.2.  
 Xcnl6-2B [{0059}],{0060}.    AG24F/AG24R.  (7D).
 Xglk76-2A [{822}], 2B [{822,0049}].  [Xglk76a-2A {822}, Xglk76b-2B {822,0049}].  pTag76  
The arm locations of Xglk76-2A,B were not reported in {822}.
 Xglk331-2B {822,0049}.    pTag331.  (4A).
The arm location of Xglk331-2B was not reported in {822}.
 Xglk370-2B {822,0049}, 2D {1034}.    pTag370.  (4A).
The arm locations of Xglk370-2B,D were not reported in {822} and {1034}.
 Xglk529-2B [{822,0049}], 2D [{822}].  [Xglk529a-2B {822,0049}, Xglk529b-2D {822}].  pTag529.  
The arm locations of Xglk529-2B,D were not reported in {822}.
 Xglk539-2B {822,0049}.    pTag539.  
The arm location of Xglk539-2B was not reported in {822}.
 Xglk592-2B {822,0049}.    pTag592.  
The arm location of Xglk592-2B was not reported in {822}.
 Xglk600-2A [{822}], 2B [{822,0049}].  [Xglk600a-2A {822}, Xglk600b-2B {822,0049}.]  pTag600.  (4A).
The arm locations of Xglkj600-2A,B were not reported in {822}.
 Xglk605-2B {822,0049}.    pTag605.  
The arm location of Xglk605-2B was not reported in {822}.
 Xglk609-2B [{822,0049}], 2D [{822}].  [Xglk609b-2B {822,0049}, Xglk609a-2D {822}.]  pTag609.  (3A).
The arm locations of Xglk609-2B,D were not reported in {822}.
 Xglk618-2B {822,0049}.    pTag618.  
The arm location of Xglk618-2B was not reported in {822}.
 Xglk632-2A [{822}], 2B [{822,0049}], 2D {0031}.  [Xglk632a-2A {822}, Xglk632b-2B {822,0049}].  pTag632.  
The arm locations of Xglk632-2A,B were not reported in {822}.
 Xglk653-2A [{822}], 2B [{822,0049}], 2D {1034}.  [Xglk653a-2A {822}, Xglk653b-2B {822,0049}].  pTag653.  
The arm locations of Xglk653-2A,B and Xglk653-2D were not reported in {822} and {1034}.
 Xglk664-2A [{822}], 2B [{822,0049}], 2D {0031}.  [Xglk664a-2A {822}, Xglk664b-2B {822,0049}].  pTag664.  
The arm locations of Xglk664-2A,B were not reported in {822}.
 Xglk699-2B {822,0049}.    pTag699.  
The arm location of Xglk699-2B was not reported in {822}.
 Xglk740-2A [{822}], 2B [{822,0049}].  [Xglk740b-2A {822}, Xglk740a-2B {822,0049}.]  pTag740.  
The arm locations of Xglk740-2A,B were not reported in {822}.
 Xgwm372-2A {0035}^2^.    WMS F372/WMS R372.  
 Xksu905(Wip)-2A [{0091}].  [Wip-2A {0091}].  5C05B11 {0094}.  
 Xkvl906(Cbp2)-2A [{0091}].  [Cbp2-2A {0091}].  pBH72-B8 {0092}.  (7B).
 Xksu908(Cbp1)-2B [{0091}].  [Cbp1-2B {0091}].  RRI 10 {0095}.  (6B).
 Xksu909(Chi1)-2B [{0091}].  [Cht1a-2B {0091}].  Chi-G11 {0096}.  (3A).
 Xksu910(Tha1)-2D [{0091}].  [Tha1-2D {0091}].  CR5 {0097}.  (4A, 6B, 7A,B,D).
 XksuF41-2A.1,.2 {0078}.    pTtksuF41.  (2B,D).
 Xmwg526-2A.1,.2 {0078}.    MWG526.  (3D).
 Xpsr924-2B {0031}.    PSR924.  (1A,B,D).
 Xpsr644-2B [{0031}].  [Xpsr644b-2B {0031}].  PSR644.  (5A).
 Xpsr1200-2A,B,D {0043}.    PSR1200.  (3A).
 Xwmc41-2D {0015}.    WMC 41F/WMC 41R {0080}.  


Group 2

Amendments:

Add:

 Xbcd18-2B.1,.2,.3 [{0074}].  [Xbcd18a,b,c-2B {0074}].  BCD18.  
 Xbcd907-2B {0074}.  [Xbcd907b-2B {0074}].  BCD907.  (3B,D, 7A).
 Xbcd1086-2B [{0074}].  [Xbcd1086b-2B {0074}].  BCD1086.  
 Xpsr129-2A [{0031}].  [Xpsr129a-2A {0031}].  PSR129.  (7A,B,D).
 Xpsr961-2B {0031}.    PSR961.  (1D, 7D).
 Xsfr4(NBS)-2A [{0031}].  [PL_AP-2A {0031}].  PL_AP.  


Group 3S

Amendments:

Add:

 Xcsu358(Pal)-3B [{0091}].  [Pal-3B {0091}].  CSU358.  
 Xgdm72-3D {0030}.    ASMS F72/ASMS R72.  
 Xglk2007(Bzb2-1)-3A,B,D [{0049}].  [Bzb2-A1,B1,D1 {0049}].  IHBP-1b(c38).  
 Xgwm685-3B {0030}.    WMS F685/WMS R685.  
 XksuA1-3B {0078}.    pTtksuA1.  (1B, 5B, 7D).
 Xmsu1-3B [{0076}].    XJ5U/XJ5R.  
 Xmsu2-3B [{0076}].    XJ26U/XJ26R.  
 Xmsu3-3B [{0076}].    XJ28U/XJ28R.  
 Xrgc970-3A,B [{0031}].  [Xrgc970b-3A, Xrgc970a-3B {0031}].  RGC970.  
 Xpsr1200-3A {0031}.    PSR1200.  (2A,B,D).
 Xpsr1327-3B [{0031}].  [Xpsr1327a-3B {0031}].  PSR1327.  (1AS,DS, 1AL, 3B, 4A,5D).
 Xsfr2(Lrk10)-3B,D [{0031}].  [Lrk10-3B,D {0031}].  Lrk10.  (1A).


Group 3L

Amendments:

Add:

 Xabcp174-3B [{0087}].    ABC174 3f/ABC174 1r.  
 Xbcdp131-3D [{0087}].    BCD131 1f*/BCD 1r.  
 Note: In order to get a 3D-specific amplicon, the BCD131 1f primer was modified by removing the 5' T and adding a C at the 3' end {0087}.
 Xbcd187-3B {0078}.    BCD187.  
 Xbcd195-3B {0078}.    BCD195.  
 Xcdo251-3B {0078}.    CDO251.  
 Xcnl2-3D [{0059}],{0060}.    AC14F/AC14R.  (7B).
 Xcnl4-3D [{0031}].  [AC29 {0031}].  AC29F/AC29R {0059}.  
 Xglk118-3A {822},{0031}.    pTag118.  
 The arm location of Xglk118-3A was not reported in {822}.
 Xglk577-3A {822},{0031}, 3B,D {1034}.    pTag577.  
 The arm location of Xglk577-3A,B,D was not reported in {822} and {1034}.
 Xglk645-3A {822},{0031}.    pTag645.  
 The arm location of Xglk645-3A was not reported in {822}.
 Xglk652-3A [{0031}], 3B [{822}^1^,{0031}], 3D {9926}^4^.  [Xglk652a-3A {0031}, Xglk652b-3B {822}].  pTag652.  (1D).
 The arm location of Xglk652-3B was not reported in {822}.
 Xglk2003(Bza1-1)-3B [{0049}].  [Bza1-B1 {0049}].  IHBP-1a(1).  (5A,D, 6AL, 6BS, 7D).
 Xglk2006(Bzb-1)-3A,B,D [{0049}].  [Bzb1-A1,B1,D1 {0049}].  IHBP-1b(c1).  
 Xgwm114-3D {9929,0039}}.    WMS F114/WMS R114.  (3B).
 The arm location of Xgwm114-3D was reported incorrectly in {9929}.
 Xgwm638-3A {0035}^2^.    WMS F638/WMS R638.  
 Xgwm674-3A {9929}^1^, {0035}^2^.    WMS F674/WMS R674.  
 The arm location of Xgwm674-3A was not reported in {9929}.
 Xgwm705-3B {0030}.    WMS F705/WMS R705.  
 Xgwm707-3D {0030}.    WMS F707/WMS R707.  
 Xgwm720-3A {0030}.    WMS F720/WMS R720.  
 Xgwm751-3A {0030}.    WMS F751/WMS R751.  
 Xgwm802-3B {0030}.    WMS F802/WMS R802.  
 Xgwm853-3B {0030}.    WMS F853/WMS R853.  
Xksu909(Chi1)-3A [{0091}].  [Cht1a-3A {0091}].  Chi-G11 {0096}.  (2B).
 Xkvl912(Prp)-3A [{0091}].  [Prp-3A {0091}].  HvPRPb {00100}.  
 Xkvl914(Glb3)-3B.1,B.2,B.3,B.4,D [{0091}].  [Glb3-3B.1,B.2,B.3,B.4,D {0091}].  pBH72-I1 {0092}.  
 Xpsr936-3A {0031}.    PSR936.  

Group 3

Amendments:

Add:

 Xglk554-3B [{0031}].  [Xglk554b-3B {0031}].  pTag554.  (2A,B,D, 5B).
 Xgwm845-3B {0030}.    WMS F845/WMS R845.  
 Xpsr919-3B [{0031}].  [Xpsr919a-3B {0031}.]  PSR919.  (2A,B,D).
 Xpsr1054-3B {0031}.    PSR1054.  

 

Group 4S (4AL:4BS:4DS)

Amendments:

Add:

 Xglk348-4B [{0031}], 4D [{822},{0031}].  [Xglk348a-4B, Xglk348b-4D {0031}, Xglk348a-4D {822}].  pTag348.  
The arm location of Xglk348-4D was not reported in {822}.
 Xgwm513-4B {9929}^1^,{0035}^2^.    WMS F513/WMS R513.  
The arm location of Xgwm513-4B was not reported in {9929}.
 Xgwm601-4A {0035}^2^.    WMS F601/WMS R601.  
 Xkvl916(Oxo)-4A,B [{0091}].  [Oxo2-4A,B {0091}].  pOXOXa {00102}.  
 Xkvl917(Tha2)-4A [{0091}].  [Tha2-4A {0091}].  pBH72-C6 {0092}.  (7A,B).
 Xkvl918(Chi2)-4A,B [{0091}].  [Cht21-4A,B {0091}].  pBH72-C4 {0092}.  

 

Group 4L (4AS:4BL:4DL)

Amendments:

Add:

 Xglk315-4A {822},{0031}.    pTag315.  
 The arm location of Xglk315-4A was not reported in {822}.
 Xglk752-4A [{822}],{0031}.  [Xglk752a-4A {822}].  pTag752.  (6B).
 The arm location of Xglk752-4A was not reported in {822}.
 Xglk2004(Bza2-1)-4A,B,D [{0049}].  [Bza2-A1,B1,D1 {0049}].  IHBP-1a(c14).  
 Xgwm192-4A,B,D [{0039}].  [Xgwm192c,a,b-4A,B,D {0039}].  WMS F192/ WMS R192.  
 Xgwm397-4A {0031}.    WMS F397/WMS R397.  


Group 5AL:4BL:4DL

Amendment:

Add:

 Xgwm6-5A [{0031}].  [Xgwm6b-5A {0031}].  WMS F6/ WMS R6.  (4B,D).
It is possible that Xgwm6-5A,4B,D form a homoeologous series.
 Xkvl920(OxoLP)-4D [{0091}].  [Oxo1-4D {0091}].  pBH6-903 {00103}.  


Group 4

Amendments:

Add:

 Xglk331-4A {0031}.    pTag331.  (2B).
 Xglk600-4A [{0031}].  [Xglk600b-4A {0031}].  pTag600.  (2A,B).
 Xgwm111-4A [{0031}].  [Xgwm111c-4A {0031}].  WMS F111/WMS R111.  (7B,D).
 Xpsr934-4A [{0031}].  [Xpsr934a-4A {0031}].  PSR934.  (2A,B,D).
 Xcsl102(NBS-LRR)-4A [{0031}].  [CD16.2-4A {0031}].  Cd16.2 {0048}.  
 Xpsr967-4B {598}.    PSR967.  (1A,B, 5A, 6AS, 6BL).
 Xwpg501(Pdi)-4A,B,D {0064}.    pTAPDI501.  (1B).



Group 5S

Amendments:

Add:

 Xcdo344-5A,B,D {0034}.    CDO344.  
 Xfba114-5B {0034}, 5D {1059,0034}.    FBA114.  
The arm location of Xfba114-5D was not reported in {1059}.
 Xbfa137-5D {1059,0034}.    FBA137.  (4A).
The arm location of Xfba137-5D was not reported in {1059}.
 Xglk163-5A {0031}.  [Xglk163a-5A {0031}].  pTag163.  (1B,D, 2D, 4D, 5BL).
 Xkvl922(Tha3)-5B,D [{0091}].  [Tha3-5,D {0091}].  pBH72-K10 {0092}.  
 Xmta9-5D {1239,0034}.    MTA9 {629}.  
The arm location of Xmta9-5D was not reported in {1239}.
 Xpsr549-5A {0031}.    PSR549.  (1AS, 1AL, 2B, 3A).
 Xpsr644-5A [{0031}].  [Xpsr644a-5A {0031}].  PSR644.  (2B).
 Xrgc3-5A {0034}.    RGC3.  
 Xrgr2104-5A,D {0034}.    RGR2105.  



Group 5L

Amendments:

Add:

 Xcdo520-5A,D {0034}.    CDO520.  
 Xcdo1088 {0068}.    CDO1088.  
 Xglk163-5B [{0031}].  [Xglk163b-5B {0031}].  pTag163.  (1B,D, 2D, 4D, 5AS).
 Xglk2003(Bza-2)-5A,D [{0049}].  [Bza1-A2,D2 {0049}].  IHBP-1a(1).  (3B, 6AL, 6BS, 7D).
 Xksu919(Lpx)-5A,B [{0091}].  [Lpx-5A,B {0091}].  6C02E12 {0094}.  (4A).
 Xksu921(Mpc1)-5A [{0091}].  [Mpc1-5A {0091}].  c1 {0094}.  (7D).
 Xksu923(Pr1)-5D [{0091}].  [Pr1-5D {0091}].  CR1 [0097}.  
 Xpsp128-5A,B,D {0086}.    PSP128F1/PSP128R1.  
 Xpsp128-5D {0086}.    PSP128F2/PSP128R2.  
 Xrgc1401-5A {0067}.    C1401.  
 Xrgr2632-5A {0067}.    R2632.  
 Xrgr2856-5A {0067}.    R2856.  
 Xrgs1912-5A {0067}.    S1912.  
 Xrz596-5B,D {0034}.    RZ596.  
 Xwg232-5A,B,D {0034}.    WG232.  (1A, 4A, 7A).

4AL:5BL:5DL

Amendments:

Add:

 Xglk558-5D [{0031}].  [Xglk558a-5D {0031}].  pTag558.  (1BL, 1DS, 1D, 2B,D, 3D, 6D, 7D).
 Xgwm118-4A,5B [{0035}]^2^.  [Xgwm118a-4A, Xgwm118b-5B {0035}]^2^.  WMS F118/WMS R118.  
 Xgwm265-4A {0035}^2^.    WMS F265/WMS R265.  (4A).

Group 5

Amendments:

Add:

 Xpgh1(ELIP)-5A,B,D [{0053}].  [Xwcr12-5A,B,D {0053}].  WCR12.  
 Xglk2001(Hst1-1)-5A,B,D [{0049}].  [Hst1-A1,B1,D1 {0049}].  ITAHISTH1.  
 Xpsr967-5A {598}.    PSR967.  (1A,B, 4B, 6AS, 6BL).
 Xrgc1329-5A {0067}.    C1329.  
 Xrgr1618-5A {0067}.    R1618.  
 Xrgc2540-5A {0067}.    C2540.  

Group 6S

Amendments:


Add:

 Xcdo365-6B {0071}.    CDO365.  
 Xcnl3-6B [{0059}],{0060}.    AC22F/AC22R.  
 Xfba381-6B,D.2 {0081}.    FBA381.  (6DL).
 Xglk172-6A [{822,0049}].  [Xglk172a-6A {822,0049}].  pTag172  (7A,B).
The arm location of Xglk172-6A was not reported in {822}.
 Xglk752-6B [{822}], {0081}.  [Xglk752b {822}].  pTag752.  (4A).
The arm location of Xglk752-6B was not reported in {822}.
 Xglk2003(Bza1-4)-6B [{0049}].  [Bza1-B4 {0049}].  IHBP-1a(1).  (3B, 5A,D, 6AL, 7D).
 Xglk2005(Bza3-1)-6A,B,D [{0049}].  [Bza3-A1,B1,D1 {0049}].  IHBP-1a(17).  
 Xgwm82-6A {0035}^2^.    WMS F82/WMS R82.  
 Xksu924(Ppo)-6A,D [{0091}].  [Ppo6A,D {0091}].   7C02D02 {0094}.  
 Xksu925(Hrp)-6A [{0091}].  [Hrp-6A {0091}].  5C05D01 {0094}.  
 Xksu926(Rip)-6D [{0091}].  [Rip-6D {0091}].  5C04F01 {0094}.  
 Xpsp551-6B {0086}.    PSP551F1/PSP551R1.  
 Xrz476-6B {0081}.    RZ476.  (7B).
 Xuta1(Psif)-6A,B,D {0081}.    p26 {999}.  
 Xuta2(Psif)-6D {0081}.    p28 {999}.  


Group 6L

Amendments:

Add:

 Xabg1-6B,D {0081}.    ABG1.  
 Xabg20-6A {282}^3^,{0081}^1^, 6B {0081}, 6D {900,0081}^1^.    ABG20 {664}.  
The arm location of Xabg20-6A,D was not reported in {282} and {900}.
 Xbcd102-6A {0081}, 6B {860,0071}.    BCD102.  (2D).
 Xbcd276-6B,D {0081}.    BCD276.  
 Xcmwg644-6A,B,D {0081}.    cMWG644.  
 Xcmwg653-6A,B,D {0081}.    cMWG653.  
A Xcmwg653-6A locus was previously mapped 1.5 cM proximal to the centromere on 6AS {9927}. It is likely that Xcmwg653-6A is, in fact, located on the long arm of 6AL {0081}.
 Xcmwg716-6D {0081}.    cMWG716.  
 Xfba1-6A {0081}.    FBA1.  (4B, 6BS,DS).
 Xfba328-6B {0081}.    FBA328.  
A Xfba328-6B locus was previously mapped close to the centromere on 6BS {900}. The precise arm location had not been confirmed using ditelosomic analysis, and this locus may, in fact, be located on 6BL.
 Xfba397-6A {900,0081}, 6B,D {0081}.    FBA397.  
 Xfbb215-6A {900}, 6D {0081}.    FBB215.  
The arm location of Xfbb215-6A was not reported in {900}.
 Xfbb283-6A {900,0081}.    FBB283.  (3B).
The arm location of Xfbb283-6A was not reported in {900}.
 Xglk259-6A {822,0049}.    pTag259.  (1D).
The arm location of Xglk259-6A was not reported in {822}.
 Xglk299-6A [{822,0049}], 6D [{822}].  [Xglk299a-6A {822,0049}, Xglk299b-6D {822}].  pTag299.  
The arm locations of Xglk299-6A,D were not reported in {822}.
 Xglk512-6A [822,0049}].  [Xglk512a-6A {822,0049}].  pTag512.  (4A).
The arm location of Xglk512-6A was not reported in {822}.
 Xglk724-6A [{822,0049}], 6B,D [{822}].  [Xglk724e-6A {822,0049}, Xglk724c,b-6B,D {822}].  pTag724.  (3A,B,D, 5A).
The arm locations of Xglk724-6A,B,D were not reported in {822}.
 Xglk756-6A [{822,0049}].  [Xglk756b-6A {822,0049}].  pTag756.  (2D, 3B, 5A,D).
The arm locations of Xglk756-6A was not reported in {822}.
 Xglk2003(Bza1-3)-6A [{0049}].  [Bza1-A3 {0049}].  IHBP-1a(1).  (3B, 5A,D, 6BS, 7D).
 Xgwm356-6A {0035}^2^.    WMS F356/WMS R356.  (2A).
 XksuD1-6B {860,0081}, 6D {448}^1,4^,{0081}^1^.    pTtksuD1.  
The arm locations of XksuD1-6B and XksuD1-6D were not reported in {860} and {448}.
 Xksu908(Cbp1)-6B [{0091}].  [Cbp1-6B {0091}].  RRI 10 {0095}.  (2B).
 Xksu910(Tha1)-6B [{0091}].  [Tha1-6B {0091}].  CR5 {0097}.  (2D, 4A, 7A,B,D).
 XksuG51-6D {448}^4^,{444,0081}^1^.      
The arm location of XksuG51-6D was not reported in {448} and {444}.
 Xmwg514-6A,D {0081}.    MWG514.  
 Xmwg872-6A,B,D {0081}.    MWG872.  
 Xmwg911-6D {0081}.    MWG911.  
 Xmwg2100-6B {0081}.    MWG2100.  
 Xpsr301-6B.2 {0081}.    PSR301.  (6AS,BS,DS).
 Xpsr904-6D {0081}.    PSR904.  (3A,D, 6AS).
 Xpsr967-6B {0081}.    PSR967.  (1A,B, 4B, 5A, 6AS).
 Xtam10-6A.2 {0081}.    TAM10.  (6AS,BS).
 Xwg405-6D {0081}.    WG405.  (2D).


Group 6

Amendments:

Add:

 Xwmc104-6B {0032}.    WMC F104/WMC R104 {0037}.  

Group 7S

Amendments:

Add:

 Xcnl1-7B [{0059}].    AC1F/AC1R.  
 Xkvl906(Cbp2)-7B [{0091}].  [Cbp2-7B {0091}].  pBH72-B8 {0092}.  (2A).
 Xkvl930(Pr1)-7B,D [{0091}].  [Pr1b-7B,D {0091}].  HvPr1b {00104}.  
 Xpsr952-7B {0031}.    PSR952.  
 Xpsr955-7B {0031}.    PSR955.  

7AS:4AL:7DS

Amendments:

Add:

 Xcnl6-7D [{0059}],{0060}.    AG24F/AG24R.  (2B).
 Xglk184-7A,D [{553}].  [TAG184-7A,D {553}].  pTag184 {822}.  
The map position of Xglk184-7D in {0031} indicated that this locus belongs to the 7AS:4AL:7DS group.
 Xksu910(Tha1)-4A [{0091}].  [Tha1-4A {0091}].  CR5 {0097}.  (2D, 6B, 7A,B,D).
 Xksu919(Lpx)-4A [{0091}].  [Lpx-4A {0091}].  6C02E12 {0094}.  (5A,B).
It is not clear whether Xksu919(Lpx)-4A belongs to the group 7AS:4AL:7DS or to the group 4AL:5AL:5BL.
 Xpsp160-7A,4A {0086}.    PSP160F1/PSP160R1.  
 Xsun1-7A,D(Wx) {0077}.    SUN 1F/ SUN 1R.  
The primers SUN1F/SUN 1R amplify across an (AT)n microsatellite at the 3'end of waxy genes.

Group 7L

Amendments:

Add:

 Xbzh232(Tha)-7A,B {[{0031}].  [Xpwir232a,b-7A,B {0031}].  pWIR232 {0061}.  
 Xcdc1(Sod1)-7A,B,D {0054}.    SOD1.1 & SOD1.2.  
 Xcnl2-7B [{0059}], {0060}.    AC14F/AC14R.  (3D).
 Xglk165-7A {0031}.    pTag165 {822}.  (5B,D).
 Xglk576-7B {0031}.    pTag576 {822}.  (7AS).
 Xglk2003(Bza1-5)-7D [{0049}].  [Bza1-D5 {0049}].  IHBP-1a(1).  (3B, 5A,D, 6AL, 6BS).
 Xgwm260-7A {0035}^2^.    WMS F260/WMS R260.  
 Xksu910(Tha1)-7A,B,D [{0091}].  [Tha1-7A,B,D {0091}].  CR5 {0097}.  (2D, 4A, 6B).
 Xkvl917(Tha2)-7A,B [{0091}].  [Tha2-7A,B {0091}].  pBH72-C6 {0092}.  (4A).
 Xkvl927(Grp94)-7A.1,A.2, B [{0091}].  [Grp94-7A.1,A.2,B {0091}].  HvGRP94 {0092}.  
 Xksu928(Chi1)-7B,D [{0091}].  [Cht1b-7B,D {0091}].  Barchi3 {0096}.  
 Xksu929(Cat)-7B [{0091}].  [Cat-7B {0091}].  5C05D01 {0094}.  
 Xmwg710-7B [{0031}].  [Xmwg710a-7B {0031}].  MWG710.  (1A,B,D, 7A,4A,7D).
 Xpsr927-7B [{1181}], {0031}.  [Xpsr927.1 {1181}].  PSR927.  (4A,D).
The arm location of Xpsr927-7B was not reported in {1181}.

Group 7

Amendments:

Add:

 Xmst101-7D {0032}.    MST F101/MST R101{0038}.  
 Xksu921(Mpc1)-7D [{0091}].  [Mpc1-7D {0091].  c1 {0094}.  (5A).


Dormancy (seed)

Add at the bottom of the section: 'Tolerance to preharvest sprouting (PHS) in the cross SPR8198 x HD2329 was shown to be associated with Xwmc104-6B and Xmst101-7D, and may thus be controlled by two genes {0032}.'

Ear emergence

QEet.ocs-4A.1 {0047}. 4AL {0047}. v: CS/CS(Kanto107 4A) mapping population
ma: Associated with Wx-B1
QEet.ocs-5A.1 {0068}. 5AL {0068}. v: CS(T. spelta 5A)/CS(Cappelle-Desprez 5A) RI mapping population {0079}.
ma: Associated with Xcdo584 and morphological locus Q {0068} .

Earliness per se

Genes for earliness per se {0023} affect aspects of developmental rate that are independent of responses to vernalization and photoperiod.
Eps-A1a {0024}. 3A {0023}; 3AL {0024}. v: Chinese Spring {0024}.
Eps-A1b {0024}. v: Timstein {0024}.

Temporary symbols:

EpsWi {0025}. 3A {0025}. v: Cheyenne*7/Wichita {0025}.
epsCnn {0025}. v: Cheyenne {0025}.
QEet.ocs-5A.2
5AL {0026}. ma: Xcdo 412-5A - Xbcd9-5A region {0026}.
{0026}.

Free-threshing habit (new section)

QFt.mgb-5A 5AL {0046}. tv: Messapia/T. dicoccoides MG4343 mapping population {0046}.
{0046}.
ma:
Associated with XksuG44-5A.
QFt.mgb-6A 6A {0046}. tv: Messapia/T. dicoccoides MG4343 mapping population {0046}.
{0046}.
ma:
Associated with Xpsr312-6A.

Frost Resistance

Add: Responses to cold exposure and their genetics are reviewed in {0020}.

Glaucousness (Waxiness/Glossiness)

Epistatic inhibitors
W1I
. ma: Xcdo456-2B - 4 cM - W1I {0001}.

Height
Reduced Height

Add to preamble for Rht-1:
The Rht-1: homoeoloci are orthologous with the D8 locus in maize and the GAI locus in Arabidopsis. They encode proteins resembling nuclear transcription factors and are involved in the sensing of gibberellin levels {0019}.
Rht-A1a {0019}. v: Chinese Spring {0019}. All wheats are assumed to be monomorphic.
Rht-B1g {0019}. v: Highbury mutants M3 103-3 and M3 103-9 {0019}. Allele Rht-B1g is a fast neutron-induced mutation of Rht-B1b and produces a tall gibberellin responsive phenotype {0019}.
Rht-D1b. ma: Xpsr1871 - 1 cM - Rht-D1b - 4 cM - Xpsr821 (PhyA) {0019}.

Add below QHt.fra-1B:

QHt.ocs-4A.1 {0047}. 4AL {0047}. v: CS/CS(Kanto107 4A) mapping population {0047}.
ma: Associated with Xpsr119-4A and Wx-B1 {0047}.
QHt.ocs-4A.2 {0047}. 4AS {0047}. v: CS/CS(Kanto107 4A) mapping population {0047}.
ma: Associated with Xbcd1738-4A and Hd {0047}.
QHt.ocs-5A.1 [{0068}]. [Qt.ocs-5A.1 {0068}]. 5AL {0068}. v: CS(T. spelta 5A)/CS(Cappelle-Desprez 5A) RI mapping population {0079}.
ma: Associated with Xcdo1088 - Xbcd9 {0068}.
This QTL coincided with a QTL for culm length, QCl.ocs-5A.1 {0068}.

Leaf Tip Necrosis

Add at the end of the section:
'QTL for leaf tip necrosis were identified in {0050} and were named according to the rules for Wheat Gene Nomenclature by the catalogue curators.'
QLtn.sfr-1B
[{0050]}. 1BS {0050}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xgwm18-1B and Xglk483-1B {0050}.
QLtn.sfr-3A
[{0050]}. 3A {0050}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr570-3A and Xpsr543-3A {0050}.
QLtn.sfr-4B.1
[{0050]}. 4B {0050}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr921-4B and Xpsr593-4B [{0050}].
QLtn.sfr-4B.2
[{0050]}. 4B {0050}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr593-4B and Xpsr112-4B [{0050}].
QLtn.sfr-4D
[{0050]}. 4DL {0050}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr302-4D and Xpsr1101-4D [{0050}].
QLtn.sfr-5A
[{0050]}. 5A {0050}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr549-5A and Xglk163-5A [{0050}].
QLtn.sfr-6A
[{0050]}. 6A {0050}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr563-6A and Xpsr966-6A [{0050}].
QLtn.sfr-7B.1
[{0050]}. 7B {0050}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr350 and Xbzh232(Tha)-7B [{0050}].
QLtn.sfr-7B.2
[{0050]}. 7B {0050}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xglk750-7B and Xmwg710-7B [{0050}].
QLtn.sfr-7D
[{0050]}. 7DS {0050}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr160-7D and Xgwm44-7D [{0050}].

Lodging (new section)

QTL for lodging were identified in {0052} and were named according to the rules for Wheat Gene Nomenclature by the catalogue curators.'

QLd.sfr-1B [{0052}]. 1BS {0052}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr949-1B and Xgwm18-1B {0052}.
This QTL coincides with QTL for reduced height, increased culm stiffness and broader leave width {0052}.
QLd.sfr-2A [{0052}]. 2AS {0052}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr958-2A and Xpsr566-2A [{0052}].
This QTL coincides with QTL for reduced height, increased culm stiffness, broader leave width, more erect growth habit, later ear emergence and increased culm thickness {0052}.
QLd.sfr-2D [{0052}]. 2D {0052}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr933-2D and Xglk529-2D [{0052}].
QLd.sfr-3A [{0052}]. 3AS {0052}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr598-3A and Xpsr570-3A {0052}.
This QTL coincides with QTL for increased culm stiffness and reduced culm thickness {0052}.
QLd.sfr-4A [{0052}]. 4AS {0052}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xgwm397-4A and Xglk315-4A {0052}.
This QTL coincides with QTL for reduced height, increased culm stiffness and more erect growth habit {0052}.
QLd.sfr-5A [{0052}]. 5AL {0052}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr918-5A and Xpsr1201-5A [{0052}].
This QTL coincides with QTL for reduced height, increased culm stiffness, reduced leave width, more erect growth habit, later ear emergence and increased culm thickness {0052}.
QLd.sfr-5B [{0052}]. 5BL {0052}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr370-5B and Xpsr580-5B [{0052}].
This QTL coincides with QTL for increased culm stiffness, broader leaf width and more erect growth habit {0052}.
QLd.sfr-6B [{0052}]. 6BL {0052}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr964-6B and Xpsr142-6B {0052}.
QLd.sfr-7B [{0052}]. 7BL {0052}. v: Forno/T. spelta var. Oberkulmer mapping population {0050}.
ma: Associated with Xpsr927-7B and Xpsr350-7B {0052}.
This QTL coincides with QTL for reduced height and later ear emergence {0052}.

Nucleolus Organizer Regions

Remove the Nor-A1 entry and replace with :
Nor-A9 {00120}. [Nor-A1{221,367,835,1012}].1AS {282,276}. v: T. spelta {221,367,835,1012}. dv: T. monococcum {658,282,276}.

Remove the Nor-A3 entry and replace with :
Nor-A10 {00120}. [Nor-A3 {1014,658}]. 5AS {282,276}. dv: T. monococcum {282,276}, T. urartu IPSR (PBI) Acc. A.


Proteins

1. Grain Protein Content

Amendments:
QGpc.ndsu-6Bb. Add '{0071}' as reference for the QTL.

Add:
QGpc.ccsu-2D {0015}. 2DL {0015}. ma: Association with microsatellite locus Xwmc41-2D accounted for 19 % of the variation in grain protein content between PH132 and WL711 {0015}.

2. Enzymes
2.4. alpha-Amylase

Add at the end of a-Amy-1 section:
'Synthesis of alpha-amylase isozymes controlled by a-Amy-1 genes on chromosomes 6A and 6D is reduced in DT6BS compared to euploid CS. This result suggests the presence of a gene(s) on the long arm of chromosome 6B, which is (are) required for GA-induced alpha-amylase synthesis in the aleurone {0072}.'

3. Endosperm Storage Proteins

3.1. Glutenins
Towards the end of the preamble, between the phrases 'A system of naming the Glu-A1-1, Glu-A1-2, Glu-B1-1, and Glu-B1-2 alleles in T. turgidum var. dicoccoides is given in {796}.' and 'Following the first listing which considers the Glu-1 set for hexaploid wheat as a single locus, there is a provisional listing based on x- and y- type glutenins. These are not referenced.', insert the following phrase, then the paragraph that follows it:

'In {00116}, a comparison between spelt wheats (T. spelta) and bread wheat has been carried out for the glutenins using a nomenclature system described in {00117}.

In {00105}, the evolution of the high molecular weight glutenin loci of the A, B, D, and G genomes of wheat has been explored; 30 partial allele sequences were compared, designated by Greek letters (a,b,g, etc.) (5 of which were cited as Schlumbaum, pers. comm.; the remaining 25 have been deposited in GenBank, accession nos. X98583-X98592, X98711-X98715 and Y12401-Y12410). These partial alleles derive from all six Glu-1-1 and Glu-1-2 loci in current-day samples taken from seven species of wheat, as well as from DNA extracted from charred grain of two samples from archaeological excavations, of 3000 and 5000 years old respectively.'

The phrase following this insert, that is, as mentioned above, 'Following the first listing which considers the Glu-1 set for hexaploid wheat as a single locus, there is a provisional listing based on x- and y- type glutenins. These are not referenced.' should now comprise a new paragraph.

At the end of the Glu-A1 section, that is, between the phrase: 'The uncertainty in numbers is due to the very similar electrophoretic mobilities of some of the subunits compared with others observed either in this study or previously' and the entry for the Glu-B1 locus, add the paragraph:

'In a study including emmer wheats (T. dicoccon) {00115}, new subunits named 1+ and 2- were found in accessions MG4378/1 and MG5380/1, respectively, and provisionally assigned to Glu-A1. Until confirmed, they are not included in the Glu-A1 list.'

At the end of the Glu-B1 section, that is, between the phrase: 'it was not conclusively clear how many of these alleles were distinct from each other, or from others previously observed.' and the entry for the Glu-D1 locus, add the paragraph:

'In a study including emmer wheats (T. dicoccon) {00115}, new subunits named 7+ (in accessions MG5400/5 and MG30835/1), 8- (in accessions MG5400/5, MG30835/1, MG5333/1 and MG5507) and 13- (in accession MG5282/2) were found and provisionally assigned to Glu-B1. Until confirmed, they are not included in the Glu-B1 list.'

In the Glu-3 (LMW glutenin) section, after the paragraph 'Multiple alleles at each of the three Glu-3 wheat loci were observed {479} and effects of allelic differences on bread making quality noted {483}.', add the paragraphs:

'A series of papers {00106, 00107, 00108, and 00109} describe considerable variation in primitive wheats not present in bread wheat (A genome species T. boeoticum, T. urartu, T. thaoudar, T. aegilopoides, T. monococcum, and D-genome species T. tauschii) for the low molecular weight subunits, sufficient to use them as a source for potentially changing flour properties in bread wheat.

In {00110}, variants for LMW glutenin subunits have been reported from study of twenty-four accessions of einkorn wheat (T. monococcum ssp. monococcum). Nine of these showed two electrophoretic bands for LMW subunits, arbitrarily designated 'a' and 'b', that appeared to be associated with good bread-making quality.

In {00111}, in a study of bread and durum wheats from Portugal, the authors used the nomenclature system described in {00113} for the LMW subunits in bread wheat, and that described in {00114} for the LMW subunits in durum wheat'

3.2. Gliadins

In the preamble section, after the 'Note' that ends with the phrase: 'and {1076} studies in T. durum (19 electrophoretic patterns, referring only to variation in the omega-gliadins, in 243 accessions).' add the following phrase:

'In {00110}, variants for w-gliadins have been reported from study of twenty-four accessions of einkorn wheat (T. monococcum ssp. monococcum). In {00111}, in a study of bread and durum wheats from Portugal, the authors used the nomenclature system described in {00112} for the w-gliadins. In {00116}, a comparison between spelt wheats (T. spelta) and bread wheat has been carried out for the gliadins using a nomenclature system described in {00118}.'

Gli-A1 [{1334},{1125}]. [Gld 1A {1415}]. 1AS {150,634,1334,1607}. s: CS*/Cheyenne {634}.v: CS {150,1334,1607}.

Delete the previous corresponding entries and substitute the following:
Gli-A1a {988}. v: CS {988}; Castan {991}; Mentana {9986}; Mara {9986}; Millewa {00119}.
Gli-A1c {988}. v: Ukrainka {988}; Gazul {9985}; Sava {994}; Hopps {00119}.
Omission confirmed; this allelic designation will be used for a new allele in the future:
Gli-A1s {9981}.

Delete the previous corresponding entry and substitute the following:
Gli-A1t {9985}. v: Jeja del País {9985}; Milturum 553 {9981}; Strela {9981}.

After Gli-A1u entry, add:
Gli-A1v {9981}. v: Japhet {9981}; Rouge de Bordeaux {9981}.
Gli-B1 [{1607},{1125}.] [Gld 1B {1243,1415}, Gld-B1,-B2,-B3,-B4,-B5,-B6 {420}]. 1B {1607},1BS{150,634}.s: CS*/Cheyenne {634}. v: CS {1607,150}.

Delete the previous corresponding entries and substitute the following:
Gli-B1d {988}. v: Chopin {991}; Dneprovskaya 521 {988}; Petrel {991}; Tiberio{9986}; Yécora {9985}; Neepawa {995}; Suneca{00119}.
Gli-B1h {988}. v: Krasnodonka {988}; Pepital {991}; Rudi{991}; Cabezorro {9985}; Tincurrin {00119}.
Gli-B1l {988}. v: Clement {991}; Damier {991}; Fiocco{9986}; Avrora {9981}; Kavkaz {9981}.
Gli-B1t {9985}. v: Jeja del País {9985}.

After Gli-B1v entry, add:
Gli-B1w {9981}. v: Ardica {9981}; Barbilla (MCB-1017) {9981}.


Gli-D1 [{121}],{1125}. [Gld 1D {1415}, Gld-D1,-D2,-D3 {420}]. 1DS{121,150,634,1334,1607}. s: CS*/Cheyenne {634}. v: CS {121,150,1334,1607}.

Delete the previous corresponding entry and substitute the following:
Gli-D1n {9981}. v: Blanquillo de Toledo (MCB-0950) {9981}.

After the final entry in the Gli-D1 list (currently Gli-D1null), and before the paragraph beginning 'Three alleles at each of the Gli-1-1 (omega gliadin) loci were noted {1358}.', add:

Note: Gli-B1l encodes secalins often associated with the T1BL·1RS translocation.

Gli-A2 [{1334}],{1125}. [Gld 6A {1415}]. 6A {1334}, 6AS {1122}. v: CS.

Delete the previous corresponding entries and substitute the following:
Gli-A2c {988}. v: Siete Cerros 66 {988}; Prinqual {991}; Loreto {9986}; Escualo {9985}; Eagle {00119}.
Gli-A2d {988}. v: Dneprovskaya 521 {988}; Mocho Sobarriba{9985}; Kenyon (biotype){995}.
Gli-A2h {988}. v: Hereward {988}; Apollo {991}; N.Strampelli {9986}; Montjuich {9985}; Basalt {9981}.
Gli-A2i {988}. v: Lesostepka 75{988}; Krasnodonka{988}.
Gli-A2m {988}. v: Marquis {988}; Rex{991}; Suneca {00119}.
Gli-A2o {988}. v: Castan {991}; Touzelle{991}; Lontra {9986}; Calatrava {9985}; Glenwari {9981}.

After Gli-A2ab entry, add:
Gli-A2ac {9981}. v: Blanquillo de Barcarrota (MCB-0893) {9981}.
Gli-A2ad {9981}. v: Hembrilla Soria (MCB-1298) {9981}.
Gli-A2ae {9981}. v: Candeal de S.Lorenzo Parrilla (MCB-0932) {9981}.
Gli-A2af {9981}. v: Barbilla de Leon (MCB-1292) {9981}.
Gli-A2ag {9981}. v: Gluclub {9981}; Tincurrin {9981}.
Gli-A2ah {9981}. v: Candeal de Nava del Rey (MCB-0892) {9981}.
Gli-A2ai {9981}. v: Blanquillo (MCB-0908) {9981}.
Gli-B2 [{1607}],{1125}. [Gld 6B {1415}]. 6B {1607}, 6BS {1122}. v: CS.

Delete the previous corresponding entries and substitute the following:
Gli-B2d {988}. v: Akmolinka 1 {988}; Tselinnaya 20 {988}; Friedland {991}; César {9981}.
Gli-B2f {988}. v: Maris Freeman {988}; Master {991}; Basalt {9981}.
Gli-B2g {988}. v: Galahad {988}; Cappelle-Desprez {991}; Capitole {991}.
Gli-B2i {988}. v: Insignia {988}; Robin {9981}.
Gli-B2n {988}. v: Solo {988}; Japhet {9981}, Rouge de Bordeaux {9981}.
Gli-B2o {988}. v: Mara {9986}; Hardi {9981}; Rivoli {991}; Pippo {9986}; Slavjanka {9981}; Odesskaya 16 {988}.
Gli-B2r {991}. v: Genial {991}; Arminda {991}; Estica {991}.
Gli-B2s {988}. v: Saratovskaya 36 {988}; Aquila {9981}.
Gli-B2ab {991}. v: Orepi {991}; Bordier {9981}.

After Gli-B2af entry, add:
Gli-B2ag {9981}. v: Jeja del Pais {9985}; Barbilla de Leon (MCB-1292) {Sp.,9981}.
Gli-B2ah {9981}. v: Rojo de Humanes (MCB-1262) {9981}; Grano de Miracolo {9981}.
Gli-B2ai {9981}. v: Blanquillo (MCB-0908) {9981}.
Gli-B2aj {9981}. v: Negrete de Málaga (MCB-1754) {9981}.
Gli-B2ak {9981}. v: HY320 {9981}; Leader {9981}.
Gli-B2al {9981}. v: Dankowska {991}.
Gli-B2am {9981}. v: TM-275 {9981}; Uralochka {9981}.
Gli-B2an {9981}. v: Eagle {9981}; Glenwari {9981}.
Gli-B2ao {9981}. v: Olympic {9981}; Mokoan {9981}.
Gli-B2ap {9981}. v: Veda {9981}; Magnif 27 {9981}.

Gli-B2aq {9981}. v: Winglen {9981}; Isis {9981}.
Gli-B2ar {9981}. v: Arbon {9981}; Roazon {9981}.
Gli-B2as {9981}. v: Strela {9981}; Sredneuralskaya {9981}.
Gli-B2at {9981}. v: Ranee {9981}; Javelin 48 {9981}.
Gli-D2 [{1334}], {1125}. [Gld 6D {1415}]. 6D {1334}, 6DS {1122}. v: CS.


Delete the previous corresponding entries and substitute the following:
Gli-D2f {988}. v: Rempart {991}; Créneau {991}; Kirgizskaya Yubileinaya {988}.
Gli-D2h {988}. v: Capitole {991}; Garant {991}; Thatcher {995}; Chinook {995}; Sadovo 1 {988}; Eagle {00119}
Gli-D2i {988}. v: Insignia 49 {00119}; Lario {9986}.
Gli-D2k {988}. v: Skala {988}; Crvencapa {994}; Kzyl-Bas {988}.

Omission confirmed; this allelic designation will be used for a new allele in the future:
Gli-D2l.

Delete the previous corresponding entries and substitute the following:
Gli-D2m {988}. v: Marquis {988}; Rex {991}; Veronese {9986}; Yecora {9985}; Rinconada {9985}; Suneca {00119}.
Gli-D2q {988}. v: Soissons {991}; Fournil {991}; E. Mottin {9981}; Volshebnitsa (biotype) {988}; Winglen {9981}; Cook {9981}.

After Glu-D2aa entry, add:
Gli-D2ab {9981}. v: Rojo de Boadilla de Campos (MCB-1031) {9981}.
Gli-D2ac {9981}. v: Albatros {9981}.
Gli-D2ad {9981}. v: Hembrilla Soria (MCB-1298) {9981}.

5. Other proteins

5.6. Waxy Proteins

Add: 'Lists of cultivars, lines and landraces of tetraploid and hexaploid wheats with different, mostly null, alleles at the Wx loci are given in {9910,9911,9912,1053,1054,9913,9915,9916,1650,9917}.'

At end of preamble add: 'All combinations of the null alleles were produced in Chinese Spring {0018}.'

Wx-A1
Add at the bottom of the section:
'The complete genomic sequence for the Wx-A1a allele from CS {0073} and the cDNA sequence for the Wx-A1b allele from Kanto 107 {0075} have been determined.'

Wx-B1
Wx-B1e
{0027}. v: Blue Boy II {0027}; Canthatch {0027}; Eureka {0027}; Götz {0027}; Norin 44 {0027}; Turkey Red {0027}.

Add at the bottom of the section:
'The complete genomic sequence for Wx-B1a from CS has been determined {0073}.'

Wx-D1
Add at the bottom of the section:
'The complete genomic sequence for Wx-D1a from CS {0073} and the cDNA sequence for the Wx-D1b allele from Bai Huo {0075} have been determined.'

5.7. Starch granule proteins

Add at the bottom of the 'Sgp-1' section: 'The proteins, designated Sgp-1, are starch synthases, encoded by SsII-A1, SsII-B1 and SsII-D1 {0042}.'

Add at the bottom of the 'Sgp-3' section: 'The proteins, designated Sgp-3, are identical to wheat starch synthase I, encoded by SsI-A1, SsI-A2 and SsI-D1 {0041}.'

5.8. Puroindolines

Pina-Am1 {0083} 5AmS {0083}. dv: T. monococcum DV92, G3116 {0083}

In T. monococcum Pina-Am1 is completely linked to Gsp-Am1 {0083}.

Pina-D1
Pina-D1a
{452}.
Replace 'carrying the Pinb-D1b mutation {452,1035}.' with 'carrying a hardness mutation in puroindoline b
{452,1035,0082}.'.
Pina-D1b {1035}.
Delete the sentence starting with 'Pinb-D1a is present ' and replace with 'Present only in some hard
hexaploid wheats.'.

Pinb-Am1 {0083} 5AmS {0083}. dv: T. monococcum DV92, G3116 {0083}

In T. monococcum Pinb-Am1 is 0.1 cM proximal to Pina-Am1 and both loci are less than 36 kb apart {0083}.

Pinb-D1
Pinb-D1a
{452}.
Add reference '{0082}.' at the end of the sentence starting with 'Pinb-D1a is present '.
Pinb-D1b {1035}.
Delete the two sentences from 'Pinb-D1b may be present ' and replace with 'Pinb-D1b is a "loss-of-function"
mutation resulting from the replacement of a glycine by a serine at position 46 {452}.'.

Pinb-D1c {0082} v: Avle {0082}; Reno {0082};Tjalve {0082}; Bjorke {0082}; Portal {0082}.
Pinb-D1c is a "loss-of-function" mutation resulting from the replacement of a leucine by a proline at position
60 {0082}.
Pinb-D1d {0082}. v: Bercy {0082}; Mjolner {0082}.
Pinb-D1d is a "loss-of-function" mutation resulting from the replacement of a tryptophan by a arginine at
position 44 {0082}.

Pinb-D1b, Pinb-D1c, or Pinb-D1d are present in hard hexaploid wheats not carrying the Pina-D1b (null) mutation {452,1035,0082}.

5.9. Starch synthase

SsI-A1 {0041}. 7A {0041}.
SsI-B1 {0041}. 7B {0041}.
SsI-D1 {0041}. 7D {0041}.
Starch synthase I proteins are identical to the starch granule proteins Sgp-3 {0041}.

SsII-A1 {0042}. 7A {0042}.
SsII-B1 {0042}. 7B {0042}.

SsII-D1 {0042}. 7D{0042}.
Starch synthase II proteins are identical to the starch granule proteins Sgp-1 {0042}.


Quality Parameters

3. Amylose content

QAmc.ocs-4A.1 {0047}. 4AS {0047}. v: CS/CS(Kanto107 4A) mapping population {0047}.
ma:
Associated with Xbcd1738 and Xcdo1387 {0047}.


Response to Vernalization

Following the first paragraph replace with:
Insensitivity is dominant.
There is an orthologous gene series on the short arms of homoeologous group 2. The "a" alleles confer the insensitive response {0063}, the contrasting allele may be referred to as "b".
Ppd-A1a {0063}. [Ppd3 {1141}]. 2AL {1268}. v: C591 {0057}.
Ppd-B1a {0063}. [Ppd2 {1566}]. 2BS {1566,1268,1269} s: Cappelle-Desprez*CS 2B {0058}. v: Chinese Spring {1268}; Spica {557}; Timstein {1269}; Sharbati Sonora Ppd-A3 {887}. ma: Xpsr666 - 1.2 cM - Xpsr109 - 4.4 cM - Ppd-B1 - 4.8 cM - Xpsr804...Cent