CATALOGUE OF GENE SYMBOLS FOR WHEAT: 2000 Supplement

R.A. McIntosh¹, K.M. Devos², J. Dubcovsky³ and W.J. Rogers⁴

¹Plant Breeding Institute, The University of Sydney, 107 Cobbitty Road, Cobbitty, N.S.W., Australia, 2570.

²John Innes Centre, Norwich Research Park, Colney, Norwich, Norfolk, NR4 7UH, U.K.

³Department of Agronomy and Range Science, University of California, Davis, 95616 CA, U.S.A.

⁴Catedra de Genetica y Fitotecnia, Universidad Nacional del Centro de la Provincia de Buenos Aires, 7300 Azul, Argentina.

The most recent edition of the Catalogue appeared in the Proceedings of the 9th International Wheat Genetics Symposium Vol. 5 (A.E. Slinkard ed., University Extension Press, University of Saskatchewan, Saskatoon, Canada). A modified version is displayed on the Graingenes Website: grains@greengenes.cit.cornell.edu

The 1999 Supplement is included in 1999 Annual Wheat Newsletter, Wheat Information Service and is listed in the Graingenes Website. The present Supplement will be offered to editors/curators for similar listing.

Revisions

10. Laboratory Designators for DNA markers

cdc	Gusta, L.V. Crop Development Centre* University of Saskatchewan 51 Campus Drive Saskatoon, Saskatchewan, S7N 5A8 Canada	ocs	Kato, K. Dept. of Crop Science* Obihiro* University of Agriculture and Veterinary Medicine Obihiro 080-8555 Japan
csl	Lagudah, E.S.* Division of Plant Industry CSIRO*, GPO Box 1600 Canberra ACT 2601 Australia	pgh	Shimosaka, E. Laboratory of Plant Genecology* Hokkaido* National Agricultural Experiment Station 1, Hitsujigaoka, Toyohira-ku Sapporo, 062-8555 Japan
kvl	Collinge, D.B. Section for Plant Pathology Dept. of Plant Biology Royal Veterinary and Agricultural University* Thorvaldsenvej 40 1871 Frederiksberg C Denmark	rgs	(Rice etiolated shoot* clones) Sasaki, T. (see rgc)
msu	Talbert, L.E. Plant Sciences Department Montana State University Bozeman, MT 59717 USA	sun	Sharp, PJ. Plant Breeding Institute University of Sydney* PMB 11, Camden NSW 2570 Australia

Morphological, Physiological, Molecular and DNA Traits

Following the first paragraph insert: More than 20 NILs involving genes affecting a range of traits are described in {0066}. These are not yet incorporated into the Catalogue.

Gross Morphology : Spike Characteristics

3. Sphaerococcum

Revision:

The naturally-occurring sphaerococcum gene in chromosome 3D and various mutant alleles conferring a similar phenotype form a homoeologous series. The sphaerococcoid alleles are either recessive or incompletely dominant. All three mapped loci are closely linked to the respective centromeres {0030}. The "a" alleles are allocated to Chinese Spring or "normal" wheats.

S-A1 {0029}.	3A {0056}.			v:	CS {0029}.
S-A1a {0029}.				v:	CS {0029}; common wheats {0029}.
S-A1b {0029}.	S3 {0056}.			v:	MS 1453 {0056}.
	ma:	Xgwm2-3A(S) - 5.1 cM - S-A1 - 6.6 cM - Xgwm720-3A(L) {0030}.
S-B1 {0029}.	3B {0030}.			v:	CS {0029}.
S-B1a {0029}.				v:	CS {0029}; common wheats {0029}.
S-B1b {0029}.	S2 {0030}.			v:	MSK 2452 {0056}; MSK 2454 {0056}.
	ma:		Xgwm685-3B(S) - 4.2 cM - S-B1 - 0.5 cM - Xgwm566/Xgwm845/cent {0030}.
S-D1 {0029}.	3D {1292,0030}; 3DS {1193,1194}; 3DL {692}.			v:	CS {0029}.
S-D1a {0029}.				v:	CS {0029}; common wheats {0029}.
S-D1b {0029}.	s1 [sp1 {1286}].			i:	S-615*11/T.sphaerococcum var. rotundatum {1500}.
				s:	CS*7/T.sphaerococcum rubiginosum 3D {1304}.
				v:	Sphaerococcum wheats {0029}.
S-D1c {0029}.	S1 {0056}.			v:	MS 3287 {0056}.
	ma:	Xgdm72-3D(S) - 8.0 cM - S-D1 - 2.9 cM - Xgwm456-3D/cent {0030}.

Temporary designation

s2 [sp2 {1286}].

Partially dominant {1286}.

Sphaerococcum simulator {1286}.

Sphaerococcum-like tetraploid wheats were reported {122,475,1282,1286}, but comparisons between them, or with s2, were not made. Whereas Schmidt & Johnson {1281} reported a single recessive factor controlling the sphaerococcum character in tetraploid wheat, Joppa {621} using the same stock found that two recessive genes were necessary to produce this phenotype.

Ear length (new section under 'Spike characteristics')

QEl.ocs-5A.1 {0068}.	5AL {0068}.		v:	CS(T. spelta 5A)/CS(Cappelle-Desprez 5A) RI mapping population {0079}.
	ma:	Associated with Xbcd9 {0068}.

Awnedness

1. Dominant Inhibitors

1.1 Hooded

Hd. Add: ‘ma: Xcdo1387-4A – 8.2 cM – Hd – 7.2 cM – Xpsr163 {0047}.’

Boron Tolerance

Bo3.	4A {0012}.

Crossability with Rye, Hordeum and Aegilops spp.

1. Common wheat

Current section

2. Tetraploid wheat

The Chinese tetraploid, Ailanmai, possesses recessive crossability genes on chromosomes 1A, 6A and 7A with the 6A gene being the least effective {0017}.

DNA Markers

Group 1S

Amendments:

Xcdo534-1B. Revise the last column to '(6A,B,D, 7A).'.

Xglk558-1D. Add ‘(5D).’ in the last column.

Xpsr549-1A.2. Add ‘(5A).’ in the last column.

Xpsr908-1B. Add '(6D).' in the last column.

Xpsr1327-1D. Revise the first column to ‘Xpsr1327-1A [{0031}], 1D {410}.’, add ‘[Xpsr1327b-1A {0031}].’ in the second column and ‘(3B).’ in the last column.

Xsfr1(Lrk10)-1A. Revise the first column to 'Xsfr2(Lrk10)-1A.', and add '(3B,D).' in the last column.

Xsfrp1(Lrk10)-1A. Revise the first column to 'Xsfrp2(Lrk10)-1A.'.

Add:

Xcnl5-1A [{0059}].		AG10F/AG10R.
Xglk301-1B [{0031}].	[Xglk301a-1B {0031}].	pTag301.	(2D, 3D, 5D, 7A,B).
Xglk317-1B [{0031}].	[Xglk317a-1B {0031}].	pTag317.	(3D, 4D, 5A, 6A).
Xgwm264-1B.1 [{9929}^1,{0003}^2].	[Xgwm264-1B {9929}, Xgwm264c-1B {0003}].	WMS F264/ WMS R264.	(1B, 3B).
Xgwm498-1A {0035}^2.		WMS F498/WMS R498.	(1BL).
Xgwm656-1A {0035}^2.		WMS F656/WMS R656.
Xkvl901(Chs)-1B,D [{0091}].	[Chs-1B,D {0091}].	pBH72-O8 {0098}.
Xkvl902(Fmt)-1B,D [{0091].	[Fmt-1B,D {0091}].	pBH72-F1 {0099}.
Xpsr593-1B [{0031}].	[Xpsr593a-1B {0031}].	PSR593.	(2B, 4B, 7B).
Xpsr642-1B {0031}.		PSR642.
Xpsr960-1B [{0031}].		PSR960.
Xsfr3(LRR)-1A,B [{0031}].	[CD9a,b-1A {0031}].	CD9.

Group 1L

Amendments:

Xbcd265-1A,B,D. Add ‘(4A).’ in the last column.

Xglk163-1B,D. Add ‘(2D, 4D, 5AS, 5BL).’ in the last column.

Xglk558-1B. Add ‘(5D).’ in the last column.

Xgwm131-1B. Revise the first column to 'Xgwm131-1B.1 [{9929,0003}].', add '[Xgwm131-1B {9929}, Xgwm131a-1B {0003}].' in the second column, and add '(1B).' in the last column.

Xgwm498-1B. Add ‘(1AS).’ in the last column.

Xmwg710-1A,B,D. Add ‘(7BL).’ in the last column.

Xpsr172(Lhcb1)-1A. Add ‘(2B).’ in the last column.

Xpsr549-1A.1. Add ‘(5A).’ in the last column.

Xpsr1327-1A. Add ‘(1AS, 3B).’ in the last column.

Add:

Xkvl903(Chi2)-1B [{0091}].	[Cht22-1B {0091}].	pBH72-N12 {0092}.
Xgwm131-1B.2 [{0003}].	[Xgwm131b-1B {0003}].	WMS F131/WMS R131.	(1B, 3B).
Xgwm636-1B [{0003}].	[Xgwm636b-1B {0003}].	WMS F636/WMS R636.	(2A).
Xpsr924-1A,B,D {0043}.		PSR924.	(2B).

Group 1

Amendments:

Xglk558-1D. Add ‘(5D).’ in the last column.

Xglk652-1D. Add ‘(3A).’ in the last column.

Xpsr386-1A. Add ‘(2A).’ in the last column.

Xwg232-1A. Add ‘(5B,D).’ in the last column.

Add:

Xgwm264-1B.2 [{0003}^2].	[Xgwm264a-1B {0003}].	WMS F264/WMS R264	(1B, 3B).
Xpsr967-1A,B {598}.		PSR967.	(4B, 5A, 6AS, 6BL).
Xwpg501(Pdi)-1B {0064}.		pTAPDI501.	(4A,B,D).

Group 2S

Amendments:

Xbcd102-2D. Add '(6A).' in the last column.

Xcdo456-2A.1,A.2,D. Revise the first column to 'Xcdo456-2A.1,A.2 {1060}, 2B [{0074}], 2D {1060}.', add '[Xcdo456b-2B {0074}].' in the second column and add as a note 'The arm location of Xcdo456-2B was not reported in {0074}.'.

Xfbb40-2B. Revise the last column to '(6A,D).'.

Xgwm210-2D. Revise the first column to ‘Xgwm210-2B {9929}, 2D {1225}.’.

Xgwm264-2B. Delete.

Xgwm636-2A. Add '(1B).' in the last column.

Xpsr549-2B. Add ‘(5A).’ in the last column.

Xpsr566-2A,D. Revise the first column to ‘Xpsr566-2A {256}, 2B [{0031}], 2D {256}.’ and add ‘[Xpsr566b-2B {0031}].’ in the second column.

Xpsr593-2B. Add ‘(1B).’ in last column.

Xpsr908-2A,D. Add '(6D).' in the last column.

Add:

Xcsu182(Sod)-2B [{0091}].	[Sod-2B {0091}].	CSU182.
Xglk197-2A {0031}^1, 2B {9926}^4.		pTag197.	(7B).
Xglk222-2A {0031}, 2D {822}.		pTag222.	(5B,D).
The arm location of Xglk222-2D was not reported in {822}.
Xglk302-2B [{822,0049}].	[Xglk302b-2B {822,0049}].	pTag302.	(4A,D).
The arm location of Xglk302-2B was not reported in {822}.
Xglk398-2B.1,B.2 [{822,0049}].	[Xglk398a,b-2B {822,0049}].	pTag398.
The arm location of Xglk398-2B.1,B.2 was not reported in {822}.
Xglk400-2B {822,0049}.		pTag400.
The arm location of Xglk400-2B was not reported in {822}.
Xglk407-2B {822,0049}.		pTag407.	(5A).
The arm location of Xglk407-2B was not reported in {822}.
Xglk471-2B [{822,0049}].	[Xglk471b-2B {822,0049}.	pTag471.
Xglk546-2B.2 [{822,0049}].	[Xglk546f-2B {822,0049}.	pTag546.	(3B, 5A, 6B, 7A, 7B).
The arm location of Xglk546-2B.2 was not reported in {822}.
Xglk661-2B [{822,0049}].	[Xglk661c-2B {822,0049}.	pTag661.	(4A,B,D).
The arm location of Xglk661-2B was not reported in {822}.
Xglk687-2B {822,0049}.		pTag687.
The arm location of Xglk687-2B was not reported in {822}.
Xglk703-2B {822,0049}.		pTag703.
The arm location of Xglk703-2B was not reported in {822}.
Xglk2002(Hst2a-1)-2A,B,D [{0049}].	[Hst2a-A1,B1,D1 {0049}].	pwcH2A-4.
Xgwm71-2B [{0035}]^2.	[Xgwm71a-2B {0035}.	WMS F71/WMS R71.	(2AS, 2A, 3D).
Xgwm122-2A {9929}^1,{0035}^2.		WMS F122/WMS R122.
The arm location of Xgwm122-2A was not reported in {9929}.
Xgwm275-2A {9929}^1,{0035}^2.		WMS F275/WMS R275.
The arm location of Xgwm275-2A was not reported in {9929}.
Xgwm448-2A {9929}^1,{0035}^2.		WMS F448/WMS R448.
The arm location of Xgwm448-2A was not reported in {9929}.
Xgwm547-2B {0035}^2.		WMS F547/WMS R547.	(3B).
Xksu904(Per2)-2A,B [{0091}].	[Per2-2A,B {0091}].	pox22.3 {0093}.
Xpsr172(Lhcb1)-2B [{0031}].	[Xpsr172-2B {0031}.	PSR172.	(1A, 5A,B,D, 7A,B,D).
Xpsr386-2A [{0031}].	[Xpsr386c-2A {0031}].	PSR386.	(1A, 3B, 5A, 7A).

Group 2L

Amendments:

Xfba111-2D. Revise the last column to '(6A,B,D).'.

Xglk554-2A,B. Revise the first column to ‘Xglk554-2A [{822}]¹,{154}¹, 2B [{822,0049}¹], 2D {9926}⁴.’, revise the second column to ‘[Xglk554a-2A {822}, Xglk554c-2B {822,0049}].’, and add ‘(3B).’ in the last column..

Xglk558-2B,D. Add ‘(5D).’ in the last column.

Xgwm265-2A. Add ‘(4A).’ in the last column.

Xgwm356-2A. Add ‘(6A).’ in the last column.

Xpsr919-2A,B,D. Add ‘(3B).’ in the last column.

Xpsr934-2A,B,D. Add ‘(4A).’ in the last column.

Add:

Xbcd377-2A {0078}.		BCD377.
Xcdc2(Sod3)-2A,B,D [{0054}].		SOD3.1 & SOD3.2.
Xcnl6-2B [{0059}],{0060}.		AG24F/AG24R.	(7D).
Xglk76-2A [{822}], 2B [{822,0049}].	[Xglk76a-2A {822}, Xglk76b-2B {822,0049}].	pTag76.
The arm locations of Xglk76-2A,B were not reported in {822}.
Xglk331-2B {822,0049}.		pTag331.	(4A).
The arm location of Xglk331-2B was not reported in {822}.
Xglk370-2B {822,0049}, 2D {1034}.		pTag370.	(4A).
The arm locations of Xglk370-2B,D were not reported in {822} and {1034}.
Xglk529-2B [{822,0049}], 2D [{822}].	[Xglk529a-2B {822,0049}, Xglk529b-2D {822}].	pTag529.
The arm locations of Xglk529-2B,D were not reported in {822}.
Xglk539-2B {822,0049}.		pTag539.
The arm location of Xglk539-2B was not reported in {822}.
Xglk592-2B {822,0049}.		pTag592.
The arm location of Xglk592-2B was not reported in {822}.
Xglk600-2A [{822}], 2B [{822,0049}].	[Xglk600a-2A {822}, Xglk600b-2B {822,0049}.	pTag600.	(4A).
The arm locations of Xglkj600-2A,B were not reported in {822}.
Xglk605-2B {822,0049}.		pTag605.
The arm location of Xglk605-2B was not reported in {822}.
Xglk609-2B [{822,0049}], 2D [{822}].	[Xglk609b-2B {822,0049}, Xglk609a-2D {822}.	pTag609.	(3A).
The arm locations of Xglk609-2B,D were not reported in {822}.
Xglk618-2B {822,0049}.		pTag618.
The arm location of Xglk618-2B was not reported in {822}.
Xglk632-2A [{822}],2B [{822,0049}], 2D {0031}.	[Xglk632a-2A {822, Xglk632b-2B {822,0049}].	pTag632.
The arm locations of Xglk632-2A,B were not reported in {822}.
Xglk653-2A [{822}], 2B [{822,0049}], 2D {1034}.	[Xglk653a-2A {822}, Xglk653b-2B {822,0049}].	pTag653.
The arm locations of Xglk653-2A,B and Xglk653-2D were not reported in {822} and {1034}.
Xglk664-2A [{822}], 2B [{822,0049}], 2D {0031}.	[Xglk664a-2A {822}, Xglk664b-2B {822,0049}].	pTag664.
The arm locations of Xglk664-2A,B were not reported in {822}.
Xglk699-2B {822,0049}.		pTag699.
The arm location of Xglk699-2B was not reported in {822}.
Xglk740-2A [{822}], 2B [{822,0049}].	[Xglk740b-2A {822}, Xglk740a-2B {822,0049}.	pTag740.
The arm locations of Xglk740-2A,B were not reported in {822}.
Xgwm372-2A {0035}^2.		WMS F372/WMS R372.
Xksu905(Wip)-2A [{0091}].	[Wip-2A {0091}].	5C05B11 {0094}.
Xkvl906(Cbp2)-2A [{0091}].	[Cbp2-2A {0091}].	pBH72-B8 {0092}.	(7B).
Xksu908(Cbp1)-2B [{0091}].	[Cbp1-2B {0091}].	RRI 10 {0095}.	(6B).
Xksu909(Chi1)-2B [{0091}].	[Cht1a-2B {0091}].	Chi-G11 {0096}.	(3A).
Xksu910(Tha1)-2D [{0091}].	[Tha1-2D {0091}].	CR5 {0097}.	(4A, 6B, 7A,B,D).
XksuF41-2A.1,.2 {0078}.		pTtksuF41.	(2B,D).
Xmwg526-2A.1,.2 {0078}.		MWG526.	(3D).
Xpsr924-2B {0031}.		PSR924.	(1A,B,D).
Xpsr644-2B [{0031}].	[Xpsr644b-2B {0031}].	PSR644.	(5A).
Xpsr1200-2A,B,D {0043}.		PSR1200.	(3A).
Xwmc41-2D {0015}.		WMC 41F/WMC 41R {0080}.

Group 2

Amendments:

Xglk76-2A,B. Delete (moved to 2L).

Xglk163-2D. Add ‘(1B, 5AS, 5BL).’ in the last column.

Xglk197-2B. Delete (moved to 2S).

Xglk222-2D. Delete (moved to 2S).

Xglk278-2D. Revise the first column to ‘Xglk278-2A.1,A.2,B [{0031}], 2D {9926}⁴.’ and add ‘[Xglk278a,b-2A, Xglk278c-2B {0031}].’ in the second column.

Xglk293-2D. Revise the first column to ‘Xpsr293-2A [{0031}], 2D {822}.’ and add ‘[Xpsr293b-2A {0031}].’ in the second column.

Xglk301-2D. Add ‘(1B, 7B)’ in the last column.

Xglk302-2B. Delete (moved to 2S).

Xglk331-2B. Delete (moved to 2L).

Xglk370-2B,D. Delete (moved to 2L.

Xglk398-2B(1),(2). Delete (moved to 2S).

Xglk400-2B. Delete (moved to 2S).

Xglk407-2B. Delete (moved to 2S).

Xglk471-2B. Delete (moved to 2S).

Xglk529-2B,D. Delete (moved to 2L).

Xglk539-2B. Delete (moved to 2L).

Xglk546-2B(1),(2). Revise the first column to ‘Xglk546-2B.1 [{822}]’ and add ‘Xglk546-2B.2 was moved to 2S’.

Xglk554-2A,B,D. Delete (moved to 2L).

Xglk578-2B. Delete (moved to 2S).

Xglk592-2B. Delete (moved to 2L).

Xglk600-2A,B. Delete (moved to 2L).

Xglk605-2B. Delete (moved to 2L).

Xglk609-2B,D. Delete (moved to 2L).

Xglk610-2A. Revise the first column to ‘Xglk610-2A {822}, 2B.1,B.2,D [{0031}].’ and add ‘[Xglk610a,b-2B, Xglk610c-2D {0031}].’ in the second column.

Xglk618-2B. Delete (moved to 2L).

Xglk632-2A,B. Delete (moved to 2L).

Xglk653-2A,B,D. Delete (moved to 2L).

Xglk661-2B. Delete (moved to 2S).

Xglk664-2A,B. Delete (moved to 2L).

Xglk684-2A. Revise the first column to ‘Xglk684-2A {822}, 2B [{0031}].’ and add ‘[Xglk684b-2B {0031}].’ in the second column.

Xglk687-2B. Revise the first column to ‘Xglk687-2A [{0031}], 2B {822}.’ and add ‘[Xglk687a-2A {0031}].’ in the second column.

Xglk699-2B. Revise the first column to ‘Xglk699-2A [{0031}], 2B {822}.’ and add ‘[Xglk699b-2A {0031}].’ in the second column.

Xglk703-2B. Delete (moved to 2S).

Xglk740-2A,B. Delete (moved to 2L).

Xgwm122-2A. Delete (moved to 2S).

Xgwm210-2D. Delete (the 1999 amendments for this entry should have been made in the 2S group).

Xgwm275-2A. Delete (moved to 2S).

Xgwm448-2A. Delete (moved to 2S).

Xwg405-2D. Add '(6D).' in the last column.

Add:

Xbcd18-2B.1,.2,.3 [{0074}].	[Xbcd18a,b,c-2B {0074}].	BCD18.
Xbcd907-2B {0074}.	[Xbcd907b-2B {0074}].	BCD907.	(3B,D, 7A).
Xbcd1086-2B [{0074}].	[Xbcd1086b-2B {0074}].	BCD1086.
Xpsr129-2A [{0031}].	[Xpsr129a-2A {0031}].	PSR129.	(7A,B,D).
Xpsr961-2B {0031}.		PSR961.
Xsfr4(NBS)-2A [{0031}].	[PL_AP-2A {0031}].	PL_AP.

Group 3S

Amendments:

Xbcd907-3B,D. Add '(2B, 7A).' in the last column.

Xgwm114-3D. Delete (moved to 3L).

Add:

Xbcd907-3B.1,.2 {0078}.		BCD907.	(2B, 3D, 7A).
Xcsu358(Pal)-3B [{0091}].	[Pal-3B {0091}].	CSU358.
Xgdm72-3D {0030}.		ASMS F72/ASMS R72.
Xglk2007(Bzb2-1)-3A,B,D [{0049}].	[Bzb2-A1,B1,D1 {0049}].	IHBP-1b(c38).
Xgwm685-3B {0030}.		WMS F685/WMS R685.
XksuA1-3B {0078}.		pTtksuA1.	(1B, 5B, 7D).
Xmsu1-3B [{0076}].		XJ5U/XJ5R.
Xmsu2-3B [{0076}].		XJ26U/XJ26R.
Xmsu3-3B [{0076}].		XJ28U/XJ28R.
Xrgc970-3A,B [{0031}].	[Xrgc970b-3A, Xrgc970a-3B {0031}].	RGC970.
Xpsr1200-3A {0031}.		PSR1200.	(2A,B,D).
Xpsr1327-3B [{0031}].	[Xpsr1327a-3B {0031}].	PSR1327.	(1AS,DS, 1AL, 3B, 4A,5D).
Xsfr2(Lrk10)-3B,D [{0031}].	[Lrk10-3B,D {0031}].	Lrk10.	(1A).

Group 3L

Amendments:

Xglk609-3A. Add ‘(2B,D).’ in the last column.

Xgwm547-3B. Add ‘(2B).’ in the last column.

XksuD19-3D. Revise the first column to 'XksuD19-3B {0078}, 3D {1061}.'

Xpsr549-3A. Add ‘(5A).’ in the last column.

Xpsr904-3A,D. Revise the last column to '(6AS, 6DL).'.

Add:

Xabcp174-3B [{0087}].		ABC174 3f/ABC174 1r.
Xbcdp131-3D [{0087}].		BCD131 1f*/BCD 1r.
Note : In order to get a 3D-specific amplicon, the BCD131 1f primer was modified by removing the 5' T and adding a C at the 3' end {0087}.
Xbcd187-3B {0078}.		BCD187.
Xbcd195-3B {0078}.		BCD195.
Xcdo251-3B {0078}.		CDO251.
Xcnl2-3D [{0059}],{0060}.		AC14F/AC14R	(7B).
Xcnl4-3D [{0031}].	[AC29 {0031}].	AC29F/AC29R {0059}.
Xglk118-3A {822},{0031}.		pTag118.
The arm location of Xglk118-3A was not reported in {822}.
Xglk577-3A {822},{0031}, 3B,D {1034}.		pTag577.
The arm location of Xglk577-3A,B,D was not reported in {822} and {1034}.
Xglk645-3A {822},{0031}.		pTag645.
The arm location of Xglk645-3A was not reported in {822}.
Xglk652-3A [{0031}], 3B [{822}^1,{0031}], 3D {9926}^4.	[Xglk652a-3A {0031}, Xglk652b-3B {822}].	pTag652.	(1D).
The arm location of Xglk652-3B was not reported in {822}.
Xglk2003(Bza1-1)-3B [{0049}].	[Bza1-B1 {0049}].	IHBP-1a(1).	(5A,D, 6AL, 6BS, 7D).
Xglk2006(Bzb-1)-3A,B,D [{0049}].	[Bzb1-A1,B1,D1 {0049}].	IHBP-1b(c1).
Xgwm114-3D {9929,0039}}.		WMS F114/WMS R114.	(3B).
The arm location of Xgwm114-3D was reported incorrectly in {9929}.
Xgwm638-3A {0035}^2.		WMS F638/WMS R638.
Xgwm674-3A {9929}^1,{0035}^2.		WMS F674/WMS R674.
The arm location of Xgwm674-3A was not reported in {9929}.
Xgwm705-3B {0030}.		WMS F705/WMS R705.
Xgwm707-3D {0030}.		WMS F707/WMS R707.
Xgwm720-3A {0030}.		WMS F720/WMS R720.
Xgwm751-3A {0030}.		WMS F751/WMS R751.
Xgwm802-3B {0030}.		WMS F802/WMS R802.
Xgwm853-3B {0030}.		WMS F853/WMS R853.
Xksu909(Chi1)-3A [{0091}].	[Cht1a-3A {0091}].	Chi-G11 {0096}.	(2B).
Xkvl912(Prp)-3A [{0091}].	[Prp-3A {0091}].	HvPRPb {00100}.
Xkvl914(Glb3)-3B.1,B.2,B.3,B.4,D [{0091}].	[Glb3-3B.1,B.2,B.3,B.4,D {0091}].	pBH72-I1 {0092}.
Xpsr936-3A {0031}.		PSR936.

Group 3

Amendments:

Xglk118-3A. Delete (moved to 3L).

Xglk301-3D. Add ‘(1B, 7B).’ in the last column.

Xglk317-3D. Add ‘(1B).’ in the last column.

Xglk558-3D. Add ‘(5D).’ in the last column.

Xglk577-3A,B,D. Delete (moved to 3L).

Xglk645-3A. Delete (moved to 3L).

Xglk652-3B. Delete (moved to 3L).

Xgwm674-3A. Delete (moved to 3L).

Xmwg526-3D. Add '(2A).' in the last column.

Xpsr386-3B. Add ‘(2A).’ in the last column.

Add:

Xglk554-3B [{0031}].	[Xglk554b-3B {0031}].	pTag554.	(2A,B,D, 5B).
Xgwm845-3B {0030}.		WMS F845/WMS R845.
Xpsr919-3B [{0031}].	[Xpsr919a-3B {0031}.	PSR919.	(2A,B,D).
Xpsr1054-3B {0031}.		PSR1054.

Group 4S (4AL:4BS:4DS)

Amendments:

Xbcd265. Revise the first column to ‘Xbcd265-4B {0047}, 4D {1059}.’

Xcdo1338-4A. Add ‘(5B,D)’ in the last column.

Xglk278-4D. Add ‘(2A,B).’ in the last column.

Xpsr593-4B. Add ‘(1B).’ in last column.

Xpsr1327-4A. Add ‘(1AS, 3B).’ in the last column.

Xpsr1871(Pki)-4A,B,D. Revise the first column to 'Xkvl1871(14-3-3a)-4A,B,D [{255},{0091].' , add '[Xpsr1871(Pki)-4A,B,D {255}, 1433a-4A,B {0091}].' in the second column, and revise the third column to 'pBT6-5tot (pHv14-3-3a)'.

Add:

Xglk348-4B [{0031}], 4D [{822},{0031}].	[Xglk348a-4B, Xglk348b-4D {0031}, Xglk348a-4D {822}].	pTag348.
The arm location of Xglk348-4D was not reported in {822}.
Xgwm513-4B {9929}^1,{0035}^2.		WMS F513/WMS R513.
The arm location of Xgwm513-4B was not reported in {9929}.
Xgwm601-4A {0035}^2.		WMS F601/WMS R601.
Xkvl916(Oxo)-4A,B [{0091}].	[Oxo2-4A,B {0091}].	pOXOXa {00102}.
Xkvl917(Tha2)-4A [{0091}].	[Tha2-4A {0091}].	pBH72-C6 {0092}.	(7A,B).
Xkvl918(Chi2)-4A,B [{0091}].	[Cht21-4A,B {0091}].	pBH72-C4 {0092}.

Group 4L (4AS:4BL:4DL)

Amendments:

Xgwm6-4B {1226}. Add ‘(5A, 4D).’ in the last column.

It is possible that Xgwm6-5A,4B,D form a homoeologous series.

Xgwm6-4D {1226}. Add ‘(5A,4B).’ in the last column.

It is possible that Xgwm6-5A,4B,D form a homoeologous series.

Add:

Xglk315-4A {822},{0031}.		pTag315.
The arm location of Xglk315-4A was not reported in {822}.
Xglk752-4A [{822}],{0031}.	[Xglk752a-4A {822}].	pTag752.	(6B).
The arm location of Xglk752-4A was not reported in {822}.
Xglk2004(Bza2-1)-4A,B,D [{0049}].	[Bza2-A1,B1,D1 {0049}].	IHBP-1a(c14).
Xgwm192-4A,B,D [{0039}].	[Xgwm192c,a,b-4A,B,D {0039}].	WMS F192/ WMS R192.
Xgwm397-4A {0031}.		WMS F397/WMS R397.

Group 5AL:4BL:4DL

Amendments:

Xfba1-4B. Revise the last column to '(6BS,DS, 6AL).'.

Add:

Xgwm6-5A [{0031}].	[Xgwm6b-5A {0031}].	WMS F6/ WMS R6.	(4B,D).
It is possible that Xgwm6-5A,4B,D form a homoeologous series.
Xkvl920(OxoLP)-4D [{0091}].	[Oxo1-4D {0091}].	pBH6-903 {00103}.

Group 4

Amendments:

Xglk163-4D. Add ‘(1B, 5AS, 5BL).’ in the last column.

Xglk302-4A. Revise the first column to ‘Xglk302-4A [{822}], 4D [{0031}].’ and add ‘[Xglk302b-4D {0031}].’ in the second column.

Xglk315-4A. Delete (moved to 4L).

Xglk317-4D. Add ‘(1B).’ in the last column.

Xglk348-4D. Delete (moved to 4S).

Xglk752-4A. Delete (moved to 4L).

Xgwm513-4B. Delete (moved to 4S).

Xwg232-4A. Add ‘(5B,D).’ in the last column.

Add:

Xglk331-4A {0031}.		pTag331.	(2B).
Xglk600-4A [{0031}].	[Xglk600b-4A {0031}].	pTag600.	(2A,B).
Xgwm111-4A [{0031}].	[Xgwm111c-4A {0031}].	WMS F111/WMS R111.	(7B,D).
Xpsr934-4A [{0031}].	[Xpsr934a-4A {0031}].	PSR934.	(2A,B,D).
Xcsl102(NBS-LRR)-4A [{0031}].	[CD16.2-4A {0031}].	Cd16.2 {0048}.
Xpsr967-4B {598}.		PSR967.	(1A,B, 5A, 6AS, 6BL).
Xwpg501(Pdi)-4A,B,D {0064}.		pTAPDI501.	(1B).

Group 5S

Amendments:

Xcdo1338-5A. Revise the first column to ‘Xcdo1338-5A {282}³, 5B,D {0034}.

Xglk317-5A.1,.2, 5A. Add ‘(1B).’ in the last column.

Xgwm192-5D. Delete.

Xpsr1327-5D. Add ‘(1AS, 3B).’ in the last column.

Add:

Xcdo344-5A,B,D {0034}.		CDO344.
Xfba114-5B {0034}, 5D {1059,0034}.		FBA114.
The arm location of Xfba114-5D was not reported in {1059}.
Xbfa137-5D {1059,0034}.		FBA137.	(4A).
The arm location of Xfba137-5D was not reported in {1059}.
Xglk163-5A {0031}.	[Xglk163a-5A {0031}].	pTag163.	(1B,D, 2D, 4D, 5BL).
Xkvl922(Tha3)-5B,D [{0091}].	[Tha3-5,D {0091}].	pBH72-K10 {0092}.
Xmta9-5D {1239,0034}.		MTA9 {629}.
The arm location of Xmta9-5D was not reported in {1239}.
Xpsr549-5A {0031}.		PSR549.	(1AS, 1AL, 2B, 3A).
Xpsr644-5A [{0031}].	[Xpsr644a-5A {0031}.	PSR644.	(2B).
Xrgc3-5A {0034}.		RGC3.
Xrgr2104-5A,D {0034}.		RGR2105.

Group 5L

Amendments:

Xbcd265-5A. Add ‘(4A).’ in the last column.

Xbcd926-5A. Revise the first column to ‘Xbcd26-5A {1059}, 5B,D {0034}.’

Xbcd1087-5D. Revise the first column to ‘Xbcd1087-5A,B {0034}, 5D {446}.’

Xcdo504-5A. Revise the first column to ‘Xcdo504-5A {419,282}³, 5B {1059}, 5D {0034}.’

Xcdo584-5B. Revise the first column to 'Xcdo584-5A {0068}, 5B {1059}.'

Xcdo590-5A. Revise the first column to ‘Xcdo590-5A {9969}, 5B,D {0034}.’

Xglk165-5B,5D.1,.2. Add ‘(7B).’ in the last column.

Xglk222-5B,D. Add ‘(2A).’ in the last column.

Xglk621-5D. Delete (moved to 4AL:5BL:5DL).

XksuA1-5B. Add '(3B).' in the last column.

Xpsr918-5D. Revise the first column to ‘Xpsr918-5A,B [{0031}], 5D {1609}.’ and add ‘[Xpsr918b-5A, Xpsr918a-5B {0031}].’ in the second column.

Xrgc711-5A. Revise the first column to ‘Xrgc711-5A {9969}, 5B,D {0034}.’

Xrz474-5A. Revise the first column to ‘Xrz474-5A {9969}, 5B,D {0034}.’

Xrgr2311-5A. Revise the first column to ‘Xrgr2311-5A {9969}, 5D {0034}.’

Xrgr3226-5A. Revise the first column to ‘Xrgr3226-5A {9969}, 5D {0034}.’

Xrgr2404-5A. Revise the first column to ‘Xrgr2404-5A {9969}, 5B,D {0034}.’

Xrgr2443-5A. Revise the first column to ‘Xrgr2443-5A {9969}, 5B,D {0034}.’

Xrz630-5A. Revise the first column to ‘Xrz630-5A {9969}, 5B,D {0034}.’

Xrz698-5A. Revise the first column to ‘Xrz698-5A {9969}, 5B,D {0034}.’

Add:

Xcdo520-5A,D {0034}.		CDO520.
Xcdo1088 {0068}.		CDO1088.
Xglk163-5B [{0031}].	[Xglk163b-5B {0031}].	pTag163.	(1B,D, 2D, 4D, 5AS).
Xglk2003(Bza-2)-5A,D [{0049}].	[Bza1-A2,D2 {0049}].	IHBP-1a(1).	(3B, 6AL, 6BS, 7D).
Xksu919(Lpx)-5A,B [{0091}].	[Lpx-5A,B {0091}].	6C02E12 {0094}.	(4A).
Xksu921(Mpc1)-5A [{0091]].	[Mpc1-5A {0091}].	c1 {0094}.	(7D).
Xksu923(Pr1)-5D [{0091}].	[Pr1-5D {0091}].	CR1 [0097}.
Xpsp128-5A,B,D {0086}.		PSP128F1/PSP128R1.
Xpsp128-5D {0086}.		PSP128F2/PSP128R2.
Xrgc1401-5A {0067}.		C1401.
Xrgr2632-5A {0067}.		R2632.
Xrgr2856-5A {0067}.		R2856.
Xrgs1912-5A {0067}.		S1912.
Xrz596-5B,D {0034}.		RZ596.
Xwg232-5A,B,D {0034}.		WG232.	(1A, 4A, 7A).

4AL:5BL:5DL

Amendments:

Xgwm637-4A. Delete ‘**’ and add ‘{0035}.’ in the first colum. Delete Note: ‘Whether Xgwm637-4A belongs to … is uncertain.’.

Add:

Xglk558-5D [{0031}].	[Xglk558a-5D {0031}].	pTag558.	(1BL, 1DS, 1D, 2B,D, 3D, 6D, 7D).
Xgwm118-4A,5B [{0035}]^2.	[Xgwm118a-4A, Xgwm118b-5B {0035}]^2.	WMS F118/WMS R118.
Xgwm265-4A {0035}^2.		WMS F265/WMS R265.	(4A).

Group 5

Amendments:

Xfba114-5D {1059}. Delete (moved to 5S).

Xfba137-5D. Delete (moved to 5S).

Xglk278-5A,B. Add ‘(2A,B).’ to the last column.

Xglk301-5D. Add ‘(1B, 7B)’ in the last column.

Xmta9-5D. Delete (moved to 5S).

Xpsr172(Lhcb1)-5A,B,D. Add ‘(2B).’ in the last column.

Xpsr386-5A. Add ‘(2A).’ in the last column.

Xwg232-5A.1,B. Add ‘(5D).’ in the last column.

Xwg232-5A.2. Add ‘(5B,D).’ in the last column.

Add:

Xpgh1(ELIP)-5A,B,D [{0053}].	[Xwcr12-5A,B,D {0053}].	WCR12.
Xglk2001(Hst1-1)-5A,B,D [{0049}].	[Hst1-A1,B1,D1 {0049}].	ITAHISTH1.
Xpsr967-5A {598}.		PSR967.	(1A,B, 4B, 6AS, 6BL).
Xrgc1329-5A {0067}.		C1329.
Xrgr1618-5A {0067}.		R1618.
Xrgc2540-5A {0067}.		C2540.

Group 6S

Amendments:

Xabc173-6A,D. Revise the first column to 'Xabc173-6A {9927}², 6B {0081}¹, 6D {900}¹.'.

Xcdo534-B,D. Revise the first column to 'Xcdo534-6A {0081}¹, 6B {860}¹,{9927}², 6D {900}¹.' and revise the last column to '(1B, 7A).'.

Xcdo1380-6B. Replace '6BS' with '6BL' in 'A 6BS Xcdo1380-6B locus was mapped in {9921}.'.

Xcmwg652-6A,B,D. Revise the first column to 'Xcmwg652-6A {900}¹, 6B {9927}²,{0081}¹, 6D {9926}⁴,{0081}¹.'.

Xcmwg653-6A. Delete (moved to 6L).

Xcmwg684-6B. Add ‘(6D).’ in the last column.

Xcmwg690-6A,B. Revise the first column to 'Xcmwg690-6A,B {9927}², 6D {0081}¹.'.

Xfba1-6D. Revise the first column to 'Xfba1-6B {0081}, 6D {900}.', and add '(6AL).' in the last column.

Xfba67-6B. Revise the first column to 'Xfba67-6A {0081}, 6B {900}, 6D {0081}.'.

Xfba85-6A,D. Revise the first column to 'Xfba85-6A {900}, 6B {0081}, 6D {900}.'.

Xfba148-6A. Revise the first column to 'Xfba148-6A {900}, 6D {0081}.'.

Xfba187-6D. Revise the first column to 'Xfba187-6B {0081}, 6D {900}.'.

Xfba328-6B. Delete (moved to 6L).

Xfbb354-6D. Revise the first column to 'Xfbb354-6A,B {0081}, 6D {900}.'.

Xfbb399-6B. Revise the first column to 'Xfbb399-6A {0081}, 6B {900}, 6D {0081}.'.

Xglk479-6B. Revise the first column to ‘Xglk479-6A {822,0049}¹, 6B {900}¹, 6D {9926}⁴.’ and add ‘The arm locations of Xglk479-6A and Xglk479-6D were not reported in {822} and {9926}.’.

Xglk537-6A. Revise the first column to ‘Xglk537-6A {822,0049}¹, {9927}².’

Xglk562-6A,B. Revise the first column to ‘Xglk562-6A {822,0049}¹, 6B {9927}².’

XksuG48-6A,B,D. Revise the first column to 'XksuG48-6A {187}², 6B {187}²,{0081}¹, 6D {448}⁴,{444,862}¹.'

Xmwg966-6A. Revise the first column to 'Xmwg966-6A {9927}², 6B {0081}¹.'.

Xpsr141(Pgk2)-6A,B,D. Revise the first column to 'Xpsr141(Pgk2)-6A,B,D [{598}]^b, 6D.1,.2 {0081}.'.

Xpsr301-6A,B,D. Revise the first column to 'Xpsr301-6A {598}, 6B.1 [{598}],{0081}, 6D {598}.', add '[Xpsr301-6B {598}]' in the second column and add '(6BL).' in the last column.

Xpsr899-6A,B,D. Delete '6B {900}' from the entry.

Xpsr904-6A. Add '(6DL).' in the last column.

Xpsr967-6A. Add '(6BL).' in the last column.

Xtam10-6A,B. Revise the first column to 'Xtam10-6A.1 [{245}]¹, 6B {245}¹,{187}².', add '[Xtam10-6A {245}].' in the second column and add '(6AL).' in the last column.

Add:

Xcdo365-6B {0071}.		CDO365.
Xcnl3-6B [{0059}],{0060}.		AC22F/AC22R.
Xfba381-6B,D.2 {0081}.		FBA381.	(6DL).
Xglk172-6A [{822,0049}].	[Xglk172a-6A {822,0049}].	pTag172.	(7A,B).
The arm location of Xglk172-6A was not reported in {822}.
Xglk752-6B [{822}],{0081}.	[Xglk752b {822}].	pTag752.	(4A).
The arm location of Xglk752-6B was not reported in {822}.
Xglk2003(Bza1-4)-6B [{0049}].	[Bza1-B4 {0049}].	IHBP-1a(1).	(3B, 5A,D, 6AL, 7D).
Xglk2005(Bza3-1)-6A,B,D [{0049}].	[Bza3-A1,B1,D1 {0049}].	IHBP-1a(17).
Xgwm82-6A {0035}^2.		WMS F82/WMS R82.
Xksu924(Ppo)-6A,D [{0091}].	[Ppo6A,D {0091}].	7C02D02 {0094}.
Xksu925(Hrp)-6A [{0091}].	[Hrp-6A {0091}].	5C05D01 {0094}.
Xksu926(Rip)-6D [{0091}].	[Rip-6D {0091}].	5C04F01 {0094}.
Xpsp551-6B {0086}.		PSP551F1/PSP551R1.
Xrz476-6B {0081}.		RZ476.	(7B).
Xuta1(Psif)-6A,B,D {0081}.		p26 {999}.
Xuta2(Psif)-6D {0081}.		p28 {999}.

Group 6L

Amendments:

Xabc163-6A. Revise the first column to 'Xabc163-6A {282}³, 6D {0081}¹.'.

Xabc175-6A,D. Revise the first column to 'Xabc175-6A {9927}²,{0081}¹, 6D {900}¹.'.

Xabg388-6A. Revise the first column to 'Xabg388-6A {282}³, 6B {0081}¹.'.

Xbcd1-6B. Revise the first column to 'Xbcd1-6A {0081}¹, 6B {187}²,{0081}¹, 6D {0081}¹.'.

Xbcd357-6D. Revise the first column to 'Xbcd357-6B {860,0071}, 6D {900}.', delete '(6B).' from the last column and add 'The arm location of Xbcd357-6B was not reported in {860}.'.

Xcdo772-6A,B. Revise the first column to 'Xcdo772-6A {900}, 6B {9921}, 6D {0081}.'.

Xcmwg674-6A. Revise the first column to 'Xcmwg674-6A {9927}²,{0081}¹, 6B,D {0081}¹.'.

Xcmwg684-6A. Revise the first column to ‘Xcmwg684-6A {282}³,{9927}²,{0081}¹, 6B.2 [{9927}²,{0081}¹], 6D [{0031}]¹.’ and revise the second column to ‘[Xcmwg684-6B {9927,0081}, Xcmwg684b-6D {0031}].’.

Xcsb112(Dhn5)-6A,B. Revise the first column to 'Xcsb112(Dhn5)-6A,B [{187}]².' and add '[Xcsb112(Dhn5-6B.2,D.2 {0081}].' to the second column.

XEsi35-6A,B,D. Revise the first column to 'Xucd109(Esi35)-6A,B,D [{278}].' and add '[XEsi35-6A,B,D {278}].' in the second column.

Xfba81-6D. Revise the first column to 'Xfba81-6B {0081}, 6D {900}.'.

Xfba111-6A,B. Revise the first column to 'Xfba111-6A,B {900}, 6D {0081}.'.

Xfba381-6D. Revise the first column to 'Xfba381-6D.1 [{900}]', add '[Xfba381-6D {900}].' in the second column, and add '(6BS,6DS).' in the last column.

Xfbb40-6A. Revise the first column to 'Xfbb40-6A {900}, 6D {0081}.'.

Xfbb57-6B. Revise the first column to 'Xfbb57-6A {0081}, 6B {900}, 6D {0081}.'.

Xfbb82-6A,B. Revise the first column to 'Xfbb82-6A,B {900}, 6D {0081}.'.

Xfbb164-6B. Revise the first column to 'Xfbb164-6A {0081}, 6B {900}.'.

Xfbb169-6B,D. Revise the first column to 'Xfbb169-6A {0081}, 6B,D {900}.'.

Xfbb170-6A. Revise the first column to 'Xfbb170-6A {900}, 6B {0081}.'.

Xfbb221-6A. Revise the first column to 'Xfbb221-6A {900}, 6B {0081}.'.

Xfbb327-6B. Revise the first column to 'Xfbb327-6B {900}, 6D {0081}.'.

Xglk334-6A. Revise the first column to ‘Xglk334-6A {882,0049}¹, 6B {9927}², 6D {0081}¹.’ and revise the note to ‘The arm location of Xglk334-6A was not reported in {822}.’.

Xfbb364-6B. Revise the first column to 'Xfbb364-6B {0081}, 6B {900}, 6D {0081}.'.

Xglk547-6A. Revise the first column to ‘Xglk547-6A {9927}², 6A.1,.2,.3 [{822,0049}]¹, 6B [{822}]².’, revise the second column to’[Xglk547a,b,d-6A.1,.2,.3 {822,0049}, Xglk547c-6B {822}].’, and add ‘The arm locations of Xglk547-6A.1,.2,.3,B were not reported in {822}.’.

Xglk705-6A,B. Revise the first column to 'Xglk705-6A {9927}²,{0081}¹, 6B {822,0081}¹.'

Xglk762-6A. Revise the first column to ‘Xglk762-6A {822,0049}¹, {9927}².’ and add ‘The arm location of Xglk762-6A was not reported in {822}.'.

Xmwg19-6A. Revise the first column to 'Xmwg19-6A {9927}²,{0081}¹, 6B,D {0081}¹.'.

Xmwg21-6A. Revise the first column to 'Xmwg21-6A {9927}², 6B {0081}¹.'.

Xmwg74-6A,B. Revise the first column to 'Xmwg74-6A {9927}², 6B {900}¹, 6D {0081}¹.'.

Xmwg798-6A,B. Revise the first column to 'Xmwg798-6A {282}³,{0081}¹, 6B {9927}²,{0081}¹, 6D {0081}¹.'.

Xmwg2029-6A,B. Revise the first column to 'Xmwg2029-6A {9927}²,{0081}¹, 6B {9927}²,{0081}¹, 6D {0081}¹.'.

Xpsr463(Prk)-6A,B,D. Add '{0081}.' as reference in the first column.

Xpsr908-6B. Revise the first column to 'Xpsr908-6B {256,598}, 6D {0081}.'.

Xuta1(Psif)-6B.1,.2. Delete.

Xwg341-6B.1,.2. Revise the first column to 'Xwg341-6B.1,.2 [{444}], 6B.3 {0081}.'

Add:

Xabg1-6B,D {0081}.		ABG1.
Xabg20-6A {282}^3,{0081}^1, 6B {0081}, 6D {900,0081}^1.		ABG20 {664}.
The arm location of Xabg20-6A,D was not reported in {282} and {900}.
Xbcd102-6A {0081}, 6B {860,0071}.		BCD102.	(2D).
Xbcd276-6B,D {0081}.		BCD276.
Xcmwg644-6A,B,D {0081}.		cMWG644.
Xcmwg653-6A,B,D {0081}.		cMWG653.
A Xcmwg653-6A locus was previously mapped 1.5 cM proximal to the centromere on 6AS {9927}. It is likely that Xcmwg653-6A is, in fact, located on the long arm of 6AL {0081}.
Xcmwg716-6D {0081}.		cMWG716.
Xfba1-6A {0081}.		FBA1.	(4B, 6BS,DS).
Xfba328-6B {0081}.		FBA328.
A Xfba328-6B locus was previously mapped close to the centromere on 6BS {900}. The precise arm location had not been confirmed using ditelosomic analysis, and this locus may, in fact, be located on 6BL.
Xfba397-6A {900,0081}, 6B,D {0081}.		FBA397.
Xfbb215-6A {900}, 6D {0081}.		FBB215.
The arm location of Xfbb215-6A was not reported in {900}.
Xfbb283-6A {900,0081}.		FBB283.	(3B).
The arm location of Xfbb283-6A was not reported in {900}.
Xglk259-6A {822,0049}.		pTag259.	(1D).
The arm location of Xglk259-6A was not reported in {822}.
Xglk299-6A [{822,0049}], 6D [{822}].	[Xglk299a-6A {822,0049}, Xglk299b-6D {822}].	pTag299.
The arm locations of Xglk299-6A,D were not reported in {822}.
Xglk512-6A [822,0049}].	[Xglk512a-6A {822,0049}].	pTag512.	(4A).
The arm location of Xglk512-6A was not reported in {822}.
Xglk724-6A [{822,0049}], 6B,D [{822}].	[Xglk724e-6A {822,0049}, Xglk724c,b-6B,D {822}].	pTag724.	(3A,B,D, 5A).
The arm locations of Xglk724-6A,B,D were not reported in {822}.
Xglk756-6A [{822,0049}].	[Xglk756b-6A {822,0049}].	pTag756.	(2D, 3B, 5A,D).
The arm locations of Xglk756-6A was not reported in {822}.
Xglk2003(Bza1-3)-6A [{0049}].	[Bza1-A3 {0049}].	IHBP-1a(1).	(3B, 5A,D, 6BS, 7D).
Xgwm356-6A {0035}^2.		WMS F356/WMS R356.	(2A).
XksuD1-6B {860,0081}, 6D {448}^1,4,{0081}^1.		pTtksuD1.
The arm locations of XksuD1-6B and XksuD1-6D were not reported in {860} and {448}.
Xksu908(Cbp1)-6B [{0091}].	[Cbp1-6B {0091}].	RRI 10 {0095}.	(2B).
Xksu910(Tha1)-6B [{0091}].	[Tha1-6B {0091}].	CR5 {0097}.	(2D, 4A, 7A,B,D).
XksuG51-6D {448}^4,{444,0081}^1.
The arm location of XksuG51-6D was not reported in {448} and {444}.
Xmwg514-6A,D {0081}.		MWG514.
Xmwg872-6A,B,D {0081}.		MWG872.
Xmwg911-6D {0081}.		MWG911.
Xmwg2100-6B {0081}.		MWG2100.
Xpsr301-6B.2 {0081}.		PSR301.	(6AS,BS,DS).
Xpsr904-6D {0081}.		PSR904.	(3A,D, 6AS).
Xpsr967-6B {0081}.		PSR967.	(1A,B, 4B, 5A, 6AS).
Xtam10-6A.2 {0081}.		TAM10.	(6AS,BS).
Xwg405-6D {0081}.		WG405.	(2D).

Group 6

Amendments:

Xabg20-6A,D. Delete (moved to 6L).

Xbcd102-6B. Delete (moved to 6L).

Xbcd357-6B. Delete (moved to 6L).

Xfba397-6A. Delete (moved to 6L).

Xfbb215-6A. Delete (moved to 6L).

Xfbb283-6A. Delete (moved to 6L).

Xglk172-6A. Delete (moved to 6S).

Xglk259-6A. Delete (moved to 6L).

Xglk299-6A,D. Delete (moved to 6L).

Xglk317-6A. Add ‘(1B).’ in the last column.

Xglk479-6A,D. Delete (moved to 6S).

Xglk512-6A. Delete (moved to 6L).

Xglk547-6A(1),(2),(3),6B. Delete (moved to 6L).

Xglk558-6D. Add ‘(5D).’ in the last column.

Xglk724-6A,B,D. Delete (moved to 6L).

Xglk752-6B. Delete (moved to 6S).

XksuD1-6B,D. Delete (moved to 6L).

Xtam10-6A. Revise the last column to '(6AS,BS, 6AL).'.

Add:

Xwmc104-6B {0032}.

WMC F104/WMC R104 {0037}.

Group 7S

Amendments:

Xcdo534-7A. Revise the last column to '(1B, 6A,B,D).'.

Xglk184-7A,D. Delete (moved to 7AS:4AL:7DS).

Xglk301-7A. Revise the first column to ‘Xglk301-7A [{553}],{822}, 7B {0031}.’, add ‘(1B).’ in the last column, and add ‘The arm location of Xglk301-7A was not reported in {822}.’

Add:

Xcnl1-7B [{0059}].		AC1F/AC1R.
Xkvl906(Cbp2)-7B [{0091}].	[Cbp2-7B {0091}].	pBH72-B8 {0092}.	(2A).
Xkvl930(Pr1)-7B,D [{0091}].	[Pr1b-7B,D {0091}].	HvPr1b {00104}.
Xpsr952-7B {0031}.		PSR952.
Xpsr955-7B {0031}.		PSR955.

7AS:4AL:7DS

Amendments:

Xbcd907-7A. Add '(2B).' in the last column.

Xglk576-7A. Add ‘(7BL).’ in the last column.

Xmwg710-7A,4A,7D. Add ‘(7BL).’ in the last column.

Xpsr573-4A. Revise the first column to ‘Xpsr573-4A {255}, 7D {0031}’

Add:

Xcnl6-7D [{0059}],{0060}.		AG24F/AG24R.	(2B).
Xglk184-7A,D [{553}].	TAG184-7A,D {553}].	pTag184 {822}.
The map position of Xglk184-7D in {0031} indicated that this locus belongs to the 7AS:4AL:7DS group.
Xksu910(Tha1)-4A [{0091}].	[Tha1-4A {0091}].	CR5 {0097}.	(2D, 6B, 7A,B,D).
Xksu919(Lpx)-4A [{0091}].	[Lpx-4A {0091}].	6C02E12 {0094}.	(5A,B).
It is not clear whether Xksu919(Lpx)-4A belongs to the group 7AS:4AL:7DS or to the group 4AL:5AL:5BL.
Xpsp160-7A,4A {0086}.		PSP160F1/PSP160R1.
Xsun1-7A,D(Wx) {0077}.		SUN 1F/ SUN 1R.
The primers SUN 1F/SUN 1R amplify across an (AT)n microsatellite at the 3'end of waxy genes.

Group 7L

Amendments:

Xglk197-7B. Add ‘(2A).’ in the last column.

XksuA1-7D. Add '(3B).' in the last column.

Xpsr129-7A,B,D. Add ‘(2A).’ in the last column.

Xpsr593-7B. Add ‘(1B).’ in last column.

Xgwm111-7D. Revise the first column to ‘Xgwm111-7B [{0031}], 7D {9929}.’, add ‘Xgwm111a-7B {0031}.’ in the second column, and add ‘(4A).’ in the last column.

Xrz476-7B. Add '(6B).' in the last column.

Add:

Xbzh232(Tha)-7A,B {[{0031}].	[Xpwir232a,b-7A,B {0031}].	pWIR232 {0061}.
Xcdc1(Sod1)-7A,B,D {0054}.		SOD1.1 & SOD1.2.
Xcnl2-7B [{0059}],{0060}.		AC14F/AC14R.	(3D).
Xglk165-7A {0031}.		pTag165 {822}.	(5B,D).
Xglk576-7B {0031}.		pTag576 {822}.	(7AS).
Xglk2003(Bza1-5)-7D [{0049}].	[Bza1-D5 {0049}].	IHBP-1a(1).	(3B, 5A,D, 6AL, 6BS).
Xgwm260-7A {0035}^2.		WMS F260/WMS R260.
Xksu910(Tha1)-7A,B,D [{0091}].	[Tha1-7A,B,D {0091}].	CR5 {0097}.	(2D, 4A, 6B).
Xkvl917(Tha2)-7A,B [{0091}].	[Tha2-7A,B {0091}].	pBH72-C6 {0092}.	(4A).
Xkvl927(Grp94)-7A.1,A.2, B [{0091}].	[Grp94-7A.1,A.2,B {0091}].	HvGRP94 {0092}.
Xksu928(Chi1)-7B,D [{0091}].	[Cht1b-7B,D {0091}].	Barchi3 {0096}.
Xksu929(Cat)-7B [{0091}].	[Cat-7B {0091}].	5C05D01 {0094}.
Xmwg710-7B [{0031}].	[Xmwg710a-7B {0031}].	MWG710.	(1A,B,D, 7A,4A,7D).
Xpsr927-7B [{1181}],{0031}.	[Xpsr927.1 {1181}].	PSR927.	(4A,D).
The arm location of Xpsr927-7B was not reported in {1181}.

Group 7

Amendments:

Xglk301-7A. Delete (moved to 7S).

Xglk558-7D. Add ‘(5D).’ in the last column.

Xglk598-7B. Delete (moved to 7L).

Xpsr172(Lhcb1)-7A,B,D. Add ‘(2B).’ in the last column.

Xpsr386-7A. Add ‘(2A).’ in the last column.

Xpsr927-7B. Delete (moved to 7L).

Xwg232-7A.1. Add ‘(5B,D).’ in the last column.

Xwg232-7A.2. Add ‘(5B,D).’ in the last column.

Add:

Xmst101-7D {0032}.		MST F101/MST R101{0038}.
Xksu921(Mpc1)-7D [{0091}].	[Mpc1-7D {0091].	c1 {0094}.	(5A).

Dormancy (seed)

Add at the bottom of the section: ‘Tolerance to preharvest sprouting (PHS) in the cross SPR8198 x HD2329 was shown to be associated with Xwmc104-6B and Xmst101-7D, and may thus be controlled by two genes {0032}.’

Ear emergence

QEet.ocs-4A.1 {0047}.	4AL {0047}.		v:	CS/CS(Kanto107 4A) mapping population.
	ma:	Associated with Wx-B1.
QEet.ocs-5A.1 {0068}.	5AL {0068}.		v:	CS(T. spelta 5A)/CS(Cappelle-Desprez 5A) RI mapping population {0079}.
	ma:	Associated with Xcdo584 and morphological locus Q {0068} .

Earliness per se

Genes for earliness per se {0023} affect aspects of developmental rate that are independent of responses to vernalization and photoperiod.

Eps-A1a {0024}.	3A {0023}; 3AL {0024}.	v:	Chinese Spring {0024}.
Eps-A1b {0024}.		v:	Timstein {0024}.

Temporary symbols:

EpsWi {0025}.	3A {0025}.	v:	Cheyenne*7/Wichita {0025}.
epsCnn {0025}.		v:	Cheyenne {0025}.
QEet.ocs-5A.2 {0026}.	5AL {0026}.	ma:		Xcdo 412-5A - Xbcd9-5A region {0026}.

Free-threshing habit (new section)

QTL loci mapped include:

QFt.mgb-5A {0046}.	5AL {0046}.		tv:	Messapia/T. dicoccoides MG4343 mapping population {0046}.
	ma:	Associated with XksuG44-5A.
QFt.mgb-6A {0046}.	6A {0046}.		tv:	Messapia/T. dicoccoides MG4343 mapping population {0046}.
	ma:	Associated with Xpsr312-6A.

Frost Resistance

Add: Responses to cold exposure and their genetics are reviewed in {0020}.

Glaucousness (Waxiness/Glossiness)

Epistatic inhibitors

W1^I.

ma:

Xcdo456 -2B - 4 cM - W1^I {0001}.

Height

Reduced Height

Add to preamble for Rht-1:

The Rht-1: homoeoloci are orthologous with the D8 locus in maize and the GAI locus in Arabidopsis. They encode proteins resembling nuclear transcription factors and are involved in the sensing of gibberellin levels {0019}.

Rht-A1a {0019}.	v:	Chinese Spring {0019}. All wheats are assumed to be monomorphic.
Rht-B1g {0019}.	v:	Highbury mutants M3 103-3 and M3 103-9 {0019}. Allele Rht-B1g is a fast neutron-induced mutation of Rht-B1b and produces a tall gibberellin responsive phenotype {0019}.
Rht-D1b.	ma:	Xpsr1871 - 1cm - Rht-D1b - 4cM - Xpsr821 (PhyA) {0019}.

Add below QHt.fra-1B:

QHt.ocs-4A.1 {0047}.	4AL {0047}.		v:	CS/CS(Kanto107 4A) mapping population {0047}.
	ma:	Associated with Xpsr119-4A and Wx-B1 {0047}.
QHt.ocs-4A.2 {0047}.	4AS {0047}.		v:	CS/CS(Kanto107 4A) mapping population {0047}.
	ma:	Associated with Xbcd1738-4A and Hd {0047}.
QHt.ocs-5A.1 [{0068}].	[Qt.ocs-5A.1 {0068}].		5AL {0068}.		v:	CS(T. spelta 5A)/CS(Cappelle-Desprez 5A) RI mapping population {0079}.
	ma:	Associated with Xcdo1088 – Xbcd9 {0068}.
This QTL coincided with a QTL for culm length, QCl.ocs-5A.1 {0068}.

Leaf Tip Necrosis

Add at the end of the section:

‘QTL for leaf tip necrosis were identified in {0050} and were named according to the rules for Wheat Gene Nomenclature by the catalogue curators.'

QLtn.sfr-1B [{0050]}.	1BS {0050}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0050}.
	ma:	Associated with Xgwm18-1B and Xglk483-1B {0050}.
QLtn.sfr-3A [{0050]}.	3A {0050}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0050}.
	ma:	Associated with Xpsr570-3A and Xpsr543-3A {0050}.
QLtn.sfr-4B.1 [{0050}].	4B {0050}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0050}.
	ma:	Associated with Xpsr921-4B and Xpsr593-4B [{0050}].
QLtn.sfr-4B.2 [{0050}].	4B {0050}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0050}.
	ma:	Associated with Xpsr593-4B and Xpsr112-4B [{0050}].
QLtn.sfr-4D [{0050}].	4DL {0050}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0050}
	ma:	Associated with Xpsr302-4D and Xpsr1101-4D [{0050}].
QLtn.sfr-5A [{0050}].	5A {0050}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0050}.
	ma:	Associated with Xpsr549-5A and Xglk163-5A [{0050}].
QLtn.sfr-6A [{0050}].	6A {0050}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0050}.
	ma:	Associated with Xpsr563-6A and Xpsr966-6A [{0050}].
QLtn.sfr-7B.1 [{0050}].	7B {0050}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0050}.
	ma:	Associated with Xpsr350 and Xbzh232(Tha)-7B [{0050}].
QLtn.sfr-7B.2 [{0050}].	7B {0050}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0050}.
	ma:	Associated with Xglk750-7B and Xmwg710-7B [{0050}].
QLtn.sfr-7D [{0050}].	7DS {0050}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0050}.
	ma:	Associated with Xpsr160-7D and Xgwm44-7D [{0050}].

Lodging (new section)

QTL for lodging were identified in {0052} and were named according to the rules for Wheat Gene Nomenclature by the catalogue curators.'

QLd.sfr-1B [{0052}].	1BS {0052}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0052}.
	ma:	Associated with Xpsr949-1B and Xgwm18-1B {0052}.
This QTL coincides with QTL for reduced height, increased culm stiffness and broader leaf width {0052}.
QLd.sfr-2A [{0052}].	2AS {0052}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0052}.
	ma:	Associated with Xpsr958-2A and Xpsr566-2A [{0052}].
This QTL coincides with QTL for reduced height, increased culm stiffness, broader leaf width, more erect growth habit, later ear emergence and increased culm thickness {0052}.
QLd.sfr-2D [{0052}].	2D {0052}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0052}.
	ma:	Associated with Xpsr933-2D and Xglk529-2D [{0052}].
QLd.sfr-3A [{0052}].	3AS {0052}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0052}.
	ma:	Associated with Xpsr598-3A and Xpsr570-3A {0052}.
This QTL coincides with QTL for increased culm stiffness and reduced culm thickness {0052}.
QLd.sfr-4A [{0052}].	4AS {0052}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0052}.
	ma:	Associated with Xgwm397-4A and Xglk315-4A {0052}.
This QTL coincides with QTL for reduced height, increased culm stiffness and more erect growth habit {0052}.
QLd.sfr-5A [{0052}].	5AL {0052}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0052}.
	ma:	Associated with Xpsr918-5A and Xpsr1201-5A [{0052}].
This QTL coincides with QTL for reduced height, increased culm stiffness, reduced leaf width, more erect growth habit, later ear emergence and increased culm thickness {0052}.
QLd.sfr-5B [{0052}].	5BL {0052}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0052}.
	ma:	Associated with Xpsr370-5B and Xpsr580-5B [{0052}].
This QTL coincides with QTL for increased culm stiffness, broader leaf width and more erect growth habit {0052}.
QLd.sfr-6B [{0052}].	6BL {0052}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0052}.
	ma:	Associated with Xpsr964-6B and Xpsr142-6B {0052}.
QLd.sfr-7B [{0052}].	7BL {0052}.		v:	Forno/T. spelta var. Oberkulmer mapping population {0052}.
	ma:	Associated with Xpsr927-7B and Xpsr350-7B {0052}.
This QTL coincides with QTL for reduced height and later ear emergence {0052}.

Nucleolus Organizer Regions

Remove the Nor-A1 entry and replace with :

Nor-A9 {00120}.

[Nor-A1 {221,367,835,1012}].

1AS {282,276}.

v: dv:

T. spelta {221,367,835,1012}. T. monococcum {658,282,276}.

Remove the Nor-A3 entry and replace with :

Nor-A10 {00120}.

[Nor-A3 {1014,658}].

5AS {282,276}.

dv:

T. monococcum {282,276}, T. urartu IPSR (PBI) Acc. A.

Proteins

1. Grain Protein Content

Amendments:

QGpc.ndsu-6Bb. Add '{0071}' as reference for the QTL.

Add:

QGpc.ccsu-2D {0015}.

2DL {0015}.

ma:

Association with microsatellite locus Xwmc41-2D accounted for 19% of the variation in grain protein content between PH132 and WL711 {0015}.

2. Enzymes

2.4. alpha-Amylase

Add at the end of a-Amy-1 section:

'Synthesis of a-amylase isozymes controlled by a-Amy-1genes on chromosomes 6A and 6D is reduced in DT6BS compared to euploid CS. This result suggests the presence of a gene(s) on the long arm of chromosome 6B, which is (are) required for GA-induced a-amylase synthesis in the aleurone {0072}.'

3. Endosperm Storage Proteins

3.1. Glutenins

Towards the end of the preamble, between the phrases 'A system of naming the Glu-A1-1, Glu-A1-2, Glu-B1-1 and Glu-B1-2 alleles in T. turgidum var. dicoccoides is given in {796}.' and 'Following the first listing which considers the Glu-1 set for hexaploid wheat as a single locus, there is a provisional listing based on x- and y- type glutenins. These are not referenced.', insert the following phrase, then the paragraph that follows it:

'In {00116}, a comparison between spelt wheats (T. spelta) and bread wheat has been carried out for the glutenins using a nomenclature system described in {00117}.

In {00105}, the evolution of the high molecular weight glutenin loci of the A, B, D and G genomes of wheat has been explored; 30 partial allele sequences were compared, designated by Greek letters (a,b,g, etc.) (5 of which were cited as Schlumbaum, pers. comm.; the remaining 25 have been deposited in GenBank, accession nos. X98583-X98592, X98711-X98715 and Y12401-Y12410). These partial alleles derive from all six Glu-1-1 and Glu-1-2 loci in current-day samples taken from seven species of wheat, as well as from DNA extracted from charred grain of two samples from archaeological excavations, of 3000 and 5000 years old respectively.’

The phrase following this insert, that is, as mentioned above, 'Following the first listing which considers the Glu-1 set for hexaploid wheat as a single locus, there is a provisional listing based on x- and y- type glutenins. These are not referenced.' should now comprise a new paragraph.

At the end of the Glu-A1 section, that is, between the phrase: ‘The uncertainty in numbers is due to the very similar electrophoretic mobilities of some of the subunits compared with others observed either in this study or previously’ and the entry for the Glu-B1 locus, add the paragraph:

‘In a study including emmer wheats (T. dicoccon) {00115}, new subunits named 1⁺ and 2^- were found in accessions MG4378/1 and MG5380/1, respectively, and provisionally assigned to Glu-A1. Until confirmed, they are not included in the Glu-A1 list.’

At the end of the Glu-B1 section, that is, between the phrase: ‘it was not conclusively clear how many of these alleles were distinct from each other, or from others previously observed.’ and the entry for the Glu-D1 locus, add the paragraph:

‘In a study including emmer wheats (T. dicoccon) {00115}, new subunits named 7⁺ (in accessions MG5400/5 and MG30835/1), 8^- (in accessions MG5400/5, MG30835/1, MG5333/1 and MG5507) and 13^- (in accession MG5282/2) were found and provisionally assigned to Glu-B1. Until confirmed, they are not included in the Glu-B1 list.’

In the Glu-3 (LMW glutenin) section, after the paragraph ‘Multiple alleles at each of the three Glu-3 wheat loci were observed {479} and effects of allelic differences on bread making quality noted {483}.’, add the paragraphs:

‘A series of papers {00106, 00107, 00108 and 00109} describe considerable variation in primitive wheats not present in bread wheat (A genome species T. boeoticum, T. urartu, T. thaoudar, T. aegilopoides, T. monococcum, and D-genome species T. tauschii) for the low molecular weight subunits, sufficient to use them as a source for potentially changing flour properties in bread wheat.

In {00110}, variants for LMW glutenin subunits have been reported from study of twenty-four accessions of einkorn wheat (T. monococcum ssp. monococcum). Nine of these showed two electrophoretic bands for LMW subunits, arbitrarily designated ‘a’ and ‘b’, that appeared to be associated with good bread-making quality.

In {00111}, in a study of bread and durum wheats from Portugal, the authors used the nomenclature system described in {00113} for the LMW subunits in bread wheat, and that described in {00114} for the LMW subunits in durum wheat’

3.2. Gliadins

In the preamble section, after the ‘Note’ that ends with the phrase: ‘and {1076} studies in T. durum (19 electrophoretic patterns, referring only to variation in the omega-gliadins, in 243 accessions).’ add the following phrase:

‘In {00110}, variants for w-gliadins have been reported from study of twenty-four accessions of einkorn wheat (T. monococcum ssp. monococcum). In {00111}, in a study of bread and durum wheats from Portugal, the authors used the nomenclature system described in {00112} for the w-gliadins. In {00116}, a comparison between spelt wheats (T. spelta) and bread wheat has been carried out for the gliadins using a nomenclature system described in {00118}.’

Gli-A1 [{1334}],{1125}.		[Gld 1A {1415}].	1AS {150,634,1334,1607}.	s: CS*/Cheyenne {634}. v: CS {150,1334,1607}.
Delete the previous corresponding entries and substitute the following:
	Gli‑A1a {988}.			v: CS {988}; Castan {991}; Mentana {9986}; Mara {9986}; Millewa {00119}.
	Gli‑A1c {988}.			v: Ukrainka {988}; Gazul {9985}; Sava {994}; Hopps {00119}.
Omission confirmed; this allelic designation will be used for a new allele in the future:
	Gli‑A1s {9981}.
Delete the previous corresponding entry and substitute the following:
	Gli‑A1t {9985}.			v: Jeja del País {9985}; Milturum 553 {9981}; Strela {9981}.
After Gli-A1u entry, add:
	Gli‑A1v {9981}.			v: Japhet {9981}; Rouge de Bordeaux {9981}.
Gli-B1 [{1607}],{1125}.		[Gld 1B {1243,1415}, Gld-B1,-B2,-B3,-B4,-B5,-B6 {420}].	1B {1607}, 1BS {150,634}.	s: CS*/Cheyenne {634}. v: CS {1607,150}.
Delete the previous corresponding entries and substitute the following:
	Gli‑B1d {988}.			v: Chopin {991}; Dneprovskaya 521 {988}; Petrel {991}; Tiberio {9986}; Yécora {9985}; Neepawa {995}; Suneca {00119}.
	Gli‑B1h {988}.			v: Krasnodonka {988}; Pepital {991}; Rudi {991}; Cabezorro {9985}; Tincurrin {00119}.
	Gli‑B1l {988}.			v: Clement {991}; Damier {991}; Fiocco {9986}; Avrora {9981}; Kavkaz {9981}.
	Gli-B1t {9985}.			v: Jeja del País {9985}.
After Gli-B1v entry, add:
	Gli-B1w {9981}.			v: Ardica {9981}; Barbilla (MCB-1017) {9981}.
Gli-D1 [{121}],{1125}.		[Gld 1D {1415}, Gld-D1,-D2,-D3 {420}].	1DS {121,150,634,1334,1607}.	s: CS*/Cheyenne {634}. v: CS {121,150,1334,1607}.
Delete the previous corresponding entry and substitute the following:
	Gli‑D1n {9981}.			v: Blanquillo de Toledo (MCB-0950) {9981}.

After the final entry in the Gli-D1 list (currently Gli-D1null), and before the paragraph beginning ‘Three alleles at each of the Gli-1-1 (omega gliadin) loci were noted {1358}.’, add:

Note: Gli-B1l encodes secalins often associated with the 1BL.1RS translocation.

Gli-A2 [{1334}],{1125}.			[Gld 6A {1415}].	6A {1334}, 6AS {1122}.	v: CS.
Delete the previous corresponding entries and substitute the following:
	Gli‑A2c {988}.				v: Siete Cerros 66 {988}; Prinqual {991}; Loreto {9986}; Escualo {9985}; Eagle {00119}.
	Gli‑A2d {988}.				v: Dneprovskaya 521 {988}; Mocho Sobarriba {9985}; Kenyon (biotype) {995}.
	Gli‑A2h {988}.				v: Hereward {988}; Apollo {991}; N. Strampelli {9986}; Montjuich {9985}; Basalt {9981}.
	Gli‑A2i {988}.				v: Lesostepka 75 {988}; Krasnodonka {988}.
	Gli‑A2m {988}.				v: Marquis {988}; Rex {991}; Suneca {00119}.
	Gli‑A2o {988}.				v: Castan {991}; Touzelle {991}; Lontra {9986}; Calatrava {9985}; Glenwari {9981}.
After Gli-A2ab entry, add:
	Gli‑A2ac {9981}.				v: Blanquillo de Barcarrota (MCB-0893) {9981}.
	Gli‑A2ad {9981}.				v: Hembrilla Soria (MCB-1298) {9981}.
	Gli‑A2ae {9981}.				v: Candeal de S.Lorenzo Parrilla (MCB-0932) {9981}.
	Gli‑A2af {9981}.				v: Barbilla de Leon (MCB-1292) {9981}.
	Gli‑A2ag {9981}.				v: Gluclub {9981}; Tincurrin {9981}.
	Gli‑A2ah {9981}.				v: Candeal de Nava del Rey (MCB-0892) {9981}.
	Gli‑A2ai {9981}.				v: Blanquillo (MCB-0908) {9981}.
Gli-B2 [{1607}],{1125}.			[Gld 6B {1415}].	6B {1607}, 6BS {1122}.	v: CS.
Delete the previous corresponding entries and substitute the following:
	Gli‑B2d {988}.				v: Akmolinka 1{988}; Tselinnaya 20 {988}; Friedland {991}; César {9981}.
	Gli‑B2f {988}.				v: Maris Freeman {988}; Master {991}; Basalt {9981}.
	Gli‑B2g {988}.				v: Galahad {988}; Cappelle-Desprez {991}; Capitole {991}.
	Gli‑B2i {988}.				v: Insignia {988}; Robin {9981}.
	Gli‑B2n {988}.				v: Solo {988}; Japhet {9981}, Rouge de Bordeaux {9981}.
	Gli‑B2o {988}.				v: Mara {9986}; Hardi {9981}; Rivoli {991}; Pippo {9986}; Slavjanka {9981}; Odesskaya 16 {988}.
	Gli‑B2r {991}.				v: Genial {991}; Arminda {991}; Estica {991}.
	Gli‑B2s {988}.				v: Saratovskaya 36 {988}; Aquila {9981}.
	Gli-B2ab {991}.				v: Orepi {991}; Bordier {9981}.
After Gli-B2af entry, add:
	Gli-B2ag {9981}.				v: Jeja del Pais {9985}; Barbilla de Leon (MCB-1292) {Sp.,9981}.
	Gli-B2ah {9981}.				v: Rojo de Humanes (MCB-1262) {9981}; Grano de Miracolo {9981}.
	Gli-B2ai {9981}.				v: Blanquillo (MCB-0908) {9981}.
	Gli-B2aj {9981}.				v: Negrete de Málaga (MCB-1754) {9981}.
	Gli-B2ak {9981}.				v: HY320 {9981}; Leader {9981}.
	Gli-B2al {9981}.				v: Dankowska {991}.
	Gli-B2am {9981}.				v: TM-275 {9981}; Uralochka {9981}.
	Gli-B2an {9981}.				v: Eagle {9981}; Glenwari {9981}.
	Gli-B2ao {9981}.				v: Olympic {9981}; Mokoan {9981}.
	Gli-B2ap {9981}.				v: Veda {9981}; Magnif 27 {9981}.
	Gli-B2aq {9981}.				v: Winglen {9981}; Isis {9981}.
	Gli-B2ar {9981}.				v: Arbon {9981}; Roazon {9981}.
	Gli-B2as {9981}.				v: Strela {9981}; Sredneuralskaya {9981}.
	Gli-B2at {9981}.				v: Ranee {9981}; Javelin 48 {9981}.
Gli-D2 [{1334}], {1125}.		[Gld 6D {1415}].		6D {1334}, 6DS {1122}.	v: CS.
Delete the previous corresponding entries and substitute the following:
	Gli‑D2f {988}.				v: Rempart {991}; Créneau {991}; Kirgizskaya Yubileinaya {988}.
	Gli‑D2h {988}.				v: Capitole {991}; Garant {991}; Thatcher {995}; Chinook {995}; Sadovo 1 {988}; Eagle {00119}
	Gli‑D2i {988}.				v: Insignia 49 {00119}; Lario {9986}.
	Gli‑D2k {988}.				v: Skala {988}; Crvencapa {994}; Kzyl-Bas {988}.
Omission confirmed; this allelic designation will be used for a new allele in the future:
	Gli‑D2l.
Delete the previous corresponding entries and substitute the following:
	Gli‑D2m {988}.				v: Marquis {988}; Rex {991}; Veronese {9986}; Yecora {9985}; Rinconada {9985}; Suneca {00119}.
	Gli‑D2q {988}.				v: Soissons {991}; Fournil {991}; E. Mottin {9981}; Volshebnitsa (biotype) {988}; Winglen {9981}; Cook {9981}.
After Glu-D2aa entry, add:
	Gli‑D2ab {9981}.				v: Rojo de Boadilla de Campos (MCB-1031) {9981}.
	Gli‑D2ac {9981}.				v: Albatros {9981}.
	Gli‑D2ad {9981}.				v: Hembrilla Soria (MCB-1298) {9981}.

5. Other proteins

5.6 Waxy proteins

At end of preamble add: 'All combinations of the null alleles were produced in Chinese Spring {0018}.'

Wx-A1

Add at the bottom of the section:

'The complete genomic sequence for the Wx-A1a allele from CS {0073} and the cDNA sequence for the Wx-A1b allele from Kanto 107 {0075} have been determined.'

Wx-B1

Wx-B1e {0027}.

v:

Blue Boy II {0027}; Canthatch {0027}; Eureka {0027}; Götz {0027}; Norin 44 {0027}; Turkey Red {0027}.

Add at the bottom of the section:

'The complete genomic sequence for Wx-B1a from CS has been determined {0073}.'

Wx-D1

Add at the bottom of the section:

'The complete genomic sequence for Wx-D1a from CS {0073} and the cDNA sequence for the Wx-D1b allele from Bai Huo {0075} have been determined.'

5.7. Starch granule proteins

Add at the bottom of the ‘Sgp-1’ section: ‘The proteins, designated Sgp-1, are starch synthases, encoded by SsII-A1, SsII-B1 and SsII-D1 {0042}.’

Add at the bottom of the ‘Sgp-3’ section: ‘The proteins, designated Sgp-3, are identical to wheat starch synthase I, encoded by SsI-A1, SsI-A2 and SsI-D1 {0041}.’

5.8. Puroindolines

Pina-A^m1 {0083}.	5A^mS {0083}.	dv:	T. monococcum DV92, G3116 {0083}
In T. monococcum Pina-A^m1 is completely linked to Gsp-A^m1 {0083}.

Pina-D1.
	Pina-D1a {452}.
	Replace 'carrying the Pinb-D1b mutation {452,1035}.' with 'carrying a hardness mutation in puroindoline b {452,1035,0082}.'.
	Pina-D1b {1035}.
	Delete the sentence starting with 'Pinb-D1a is present …' and replace with 'Present only in some hard hexaploid wheats.'.

Pinb-A^m1 {0083}.	5A^mS {0083}.	dv:	T. monococcum DV92, G3116 {0083}
In T. monococcum Pinb-A^m1 is 0.1 cM proximal to Pina-A^m1 and both loci are less than 36 kb apart {0083}.

Pinb-D1.
	Pinb-D1a {452}.
	Add reference '{0082}.' at the end of the sentence starting with 'Pinb-D1a is present …'.
	Pinb-D1b {1035}.
	Delete the two sentences from 'Pinb-D1b may be present …' and replace with 'Pinb-D1b is a "loss-of-function" mutation resulting from the replacement of a glycine by a serine at position 46 {452}.'.
	Pinb-D1c {0082}.	v:	Avle {0082}; Reno {0082};Tjalve {0082}; Bjorke {0082}; Portal {0082}.
	Pinb-D1c is a "loss-of-function" mutation resulting from the replacement of a leucine by a proline at position 60 {0082}.
	Pinb-D1d {0082}.	v:	Bercy {0082}; Mjolner {0082}.
	Pinb-D1d is a "loss-of-function" mutation resulting from the replacement of a tryptophan by a arginine at position 44 {0082}.

Pinb-D1b, Pinb-D1c, or Pinb-D1d are present in hard hexaploid wheats not carrying the Pina-D1b (null) mutation {452,1035,0082}.

5.9. Starch synthase

SsI-A1 {0041}.	7A {0041}.
SsI-B1 {0041}.	7B {0041}.
SsI-D1 {0041}.	7D {0041}.

Starch synthase I proteins are identical to the starch granule proteins Sgp-3 {0041}.

SsII-A1 {0042}.	7A {0042}.
SsII-B1 {0042}.	7B {0042}.
SsII-D1 {0042}.	7D{0042}.

Starch synthase II proteins are identical to the starch granule proteins Sgp-1 {0042}.

Quality Parameters

3. Amylose content

QAmc.ocs-4A.1 {0047}.	4AS {0047}.		v:	CS/CS(Kanto107 4A) mapping population {0047}.
	ma:	Associated with Xbcd1738 and Xcdo1387 {0047}.

Response to Photoperiod

Following the first paragraph replace with:

Insensitivity is dominant.

There is an orthologous gene series on the short arms of homoeologous group 2. The "a" alleles confer the insensitive response {0063}, the contrasting allele may be referred to as "b".

Ppd-A1a {0063}.		[Ppd3 {1141}].		2AL {1268}.	v:	C591 {0057}.
Ppd-B1a {0063}.		[Ppd2 {1566}].		2BS {1566,1268,1269}	s:	Cappelle-Desprez*/CS 2B {0058}.
		v:	Chinese Spring {1268}; Spica {557}; Timstein {1269}. Sharbati Sonora Ppd-A3 {887}.
		ma:	Xpsr666 - 1.2cM - Xpsr109 - 4.4cM - Ppd-B1 - 4.8cM - Xpsr804 ….Cent {0062}.
Ppd-D1a {0063}.		[Ppd1 {1566}].		2DS {1268} [2DL pre-new arm nomenclature {1328}].
	s:		Cappelle Desprez*/Ciano 2D {1598}; Cappelle-Desprez*/Mara 2D {1598}. CS*/Ciano 2D Ppd-B1 {1268}.
		v:	Akakomugi {1604}; Bezostaya 1 {1604}; Mara {1604}; Sava {1604}; Sonora 64 {1566}. Sharbati Sonora Ppd-D1 {887}.

Two genes control photoperiod response in T. turgidum {788}.

Gene Ppd-H2 on barley chromosome 2HS may be a member of the Ppd-1 orthologous series {766}.

Response to Salinity

K⁺/Na⁺ discrimination

Add at the end of the 1^st sentence: 'Lophopyrum elongatum chromosome arms 1ES, 7ES, and 7EL enhance K⁺/Na⁺ selectivity in wheat under salt stress {0065}.

Tenacious Glumes

Tg2 {0046}.	2BS {0046}.		Derived from T. dicoccoides
	ma:	Tg2 is associated with Xrsq805(Embp)-2B and Xpsr899-2B {0046}.

Yield Components (new section)

50-grain weight

QFgw.ocs-4A.1 {0047}.	4A {0047}.		v:	CS/CS(Kanto107 4A) mapping population {0047}.
	ma:	Associated with Xbcd265-4A and Xbcd1738-4A {0047}.

Grain weight/ear

QGwe.ocs-4A.1 {0047}.	4AS {0047}.		v:	CS/CS(Kanto107 4A) mapping population {0047}.
	ma:	Associated with Xbcd1738-4A {0047}.

Plant yield

QYld.ocs-4A.1 {0047}.	4AS {0047}.		v:	CS/CS(Kanto107 4A) mapping population {0047}.
	ma:	Associated with Xbcd1738-4A {0047}.

Spikelet number/ear

QSpn.ocs-4A.1 {0047}.	4AS {0047}.		v:	CS/CS(Kanto107 4A) mapping population {0047}.
	ma:	Associated with Xbcd1738-4A {0047}.

Tiller number/plant

QTn.ocs-4A.1 {0047}.	4AS {0047}.		v:	CS/CS(Kanto107 4A) mapping population {0047}.
	ma:	Associated with Xpsr163-4A {0047}.

Pathogenic Disease/Pest Reaction

Reaction to Diuraphis noxia

Dn1	v:	Betta DN {0004}; Caledon {0004}; Gariep {0004}; Limpopo DN {0004}; Tugela DN {0004}.
Dn5	v:	Palmiet DN {0004}.

Reaction to Erysiphe graminis

Pm1.				ma:	AFLP marker 18M1 - various Pm1 alleles 0.9 cM {0011}.
Pm1c.				v:	Blaukorn {0011}. ma: AFLP marker 18M2 was diagnostic for Pm1c {0011}.
Pm3d.				v:	Kanzler {0011}.
Pm3g {0070}.	[Mlar {854}].		1A {0070}.
				ma:	Pm3g - 5.2 cM - Gli-A5 - 1.9 cM - Gli-A1 {0070}.
Pm4a.				ma:	Pm4a - 3.5 cM - AFLP markers 4aM1 and 4aM2 {0011}. Xbcd1231 was converted to a STS marker {0069}.
Pm4b.				v:	Atlantis {0011}; Boheme {0011}; Renan {0016}. RE714 {1220}.
Pm5.				v:	Greif Pm6 {0011}.
Pm6.				i:	CI 13250/7*Prins {0069}; CI 12559/8*Prins {0069}; PI 170914/7*Prins {0069}.
				v:	Greif Pm5 {0011}.
				ma:	close linkage with Xbcd135 (1.5+1.4cM), Xbcd307 (4.7+2.5cM) and Xbcd266 (4.5+2.4cM) {0069}.
Pm8.		Add just before 'crosses between three lines …': 1BS/1RS recombinants 2.9 cM proximal to Gli-B1/GluB3 {0084}. i: MA1 and MA2 four-breakpoint double translocation lines 1RS-1BS-1RS-1BS.1BL in Pavon {0084}.'
Pm13.		3B. 3D.		v: v:	add: R1B {0055}; R4A {0055}; R6A {0055}. add: R2A {0055}; R2B {0055}.
				Add at the end of the section: ‘ma: Pm13 was mapped to a translocated 3S^lS segment distal to Xcdo460-3B {0036}.
Pm17.			Add:	v:	TAM202 {0021}; Niobrara {0021}; Nekota {0021}.
Pm21.			Add:	ma:	RAPD OPH171400 and SCAR markers SCAR1400 and SCAR1265 associated with Pm21 are described in {0014}.
Pm26 {0001}.				Recessive {0001}. 2BS {0001}.
				s:	Bethlehem*8/T. turgidum var. dicoccoides 2BS {0001}. tv: T. turgidum var. dicoccoides TTD140 {0001}. ma: Co-segregation with Xwg516 {0001}.
Pm27 {0002}.		6B (6B-6G) {0002}.		v:	Line 146-155-T {0002}.
				tv:	T. timopheevii var. timopheevii K-38555 {0022}.
				ma:	6BS……Xpsr8/Xpsr964 - Pm27 - Xpsr154/Xpsr546 ……6BL {0002}. Co-segregation with Xpsr3131 {0002}.
Pm28 {0022}.		1B {0022}.		v:	Meri {0022}.
Mlre.		RE714 Pm4b		tv:	T. dicoccum 119 {1220}.

Add: Mlre showed a residual effect on the quantitative expression of APR in the presence of E. graminis pathotypes considered virulent for Mlre in standard seedling tests {0016}.

Add: List in {0028} (Finnish wheats).

Add at the end of the section:

'QTL for resistance to powdery mildew were identified in {0051} and were named according to the rules for Wheat Gene Nomenclature by the catalogue curators.'

QPm.sfr-1A [{0051}].	1A {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Oberkulmer {0051}.
	ma:	Associated with Xpsr1201-1A and Xpsr941-1A [{0051}].
QPm.sfr-1B [{0051}].	1B {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Forno {0051}.
	ma:	Associated with Xsfr3(LRR)-1B and Xpsr593-1B [{0051}].
QPm.sfr-1D [{0051}].	1D {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Oberkulmer {0051}.
	ma:	Associated with Xpsr168-1D and Xglk558-1D [{0051}].
QPm.sfr-2A [{0051}].	2A {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Oberkulmer {0051}.
	ma:	Associated with Xpsr380-2A and Xglk293-2A [{0051}].
QPm.sfr-2D [{0051}].	2D {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Oberkulmer {0051}.
	ma:	Associated with Xpsr932-2D and Xpsr331-2D [{0051}].
QPm.sfr-3A [{0051}].	3A {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Forno {0051}.
	ma:	Associated with Xpsr598-3A and Xpsr570-3A {0051}.
QPm.sfr-3D [{0051}].	3D {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Oberkulmer {0051}.
	ma:	Associated with Xpsr1196-3D and Xsfr2(Lrk10)-3D [{0051}].
QPm.sfr-4A.1 [{0051}].	4A {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Forno {0051}.
	ma:	Associated with Xgwm111-4A and Xpsr934-4A [{0051}].
QPm.sfr-4A.2 [{0051}].	4A {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Forno {0051}.
	ma:	Associated with Xmwg710-4A and Xglk128-4A [{0051}].
QPm.sfr-4B [{0051}].	4B {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Forno {0051}.
	ma:	Associated with Xpsr593-4B and Xpsr1112-4B [{0051}].
QPm.sfr-4D [{0051}].	4D {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Forno{0051}.
	ma:	Associated with Xglk302-4D and Xpsr1101-4D [{0051}].
QPm.sfr-5A.1 [{0051}].	5A {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Oberkulmer {0051}.
	ma:	Associated with Xpsr644-5A and Xpsr945-5A [{0051}].
QPm.sfr-5A.2 [{0051}].	5A {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Oberkulmer {0051}.
	ma:	Associated with Xpsr1194-5A and Xpsr918-5A [{0051}].
QPm.sfr-5B [{0051}].	5B {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Oberkulmer {0051}.
	ma:	Associated with Xpsr580-5B and Xpsr143-5B [{0051}].
QPm.sfr-6B [{0051}].	6B {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Forno {0051}.
	ma:	Associated with Xpsr167-6B and Xpsr964-6B [{0051}].
QPm.sfr-7B.1 [{0051}].	7B {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Forno {0051}.
	ma:	Associated with Xpsr593-7B and Xpsr129-7B [{0051}].
QPm.sfr-7B.2 [{0051}].	7B {0051}.		v:	Forno/T. spelta var. Oberkulmer mapping population; the resistance was contributed by Forno {0051}.
	ma:	Associated with Xglk750-7B and Xmwg710-7B [{0051}].’
	This QTL corresponds to Pm5 {0051}.

Reaction to Fusarium graminearum

QTL for fusarium head blight were identified in {0078}.