5. Other proteins

5.1. Lipopurothionins
 
Pur-A1 {351}.   1AL {351} v: CS {351}.
Pur-B1 {351}.   1BL {351}. v: CS {351}.
Pur-D1 {351}.   1DL {351}. v: CS {351}.

A locus in chromosome 5DS affects the level of lipopurothionin {351}.
 
Pur-R1.   1RL {1261}. ad: CS/Imperial.
su: Several 1R{1B} lines.
tr: Aurora, Kavkaz, with 1BL.1RS.

5.2. Lectins
 
Lec-A1.   1AL {1427}. v: CS.
Lec-B1.   1B {1427}. s: CS*/Hope 1B.
Lec-D1.   1DL {1427}. v: CS.
Lec-U1.   1U {1427}. ad: CS/Ae. umbellulata.

5.3. Iodine binding factor

A monomeric water soluble protein from mature grain which preferentially binds iodine {818}.
 
Ibf-A1 {818}.   5AL {818}. v: CS.
Ibf-A1a {818}.
    v: CS.
Ibf-A1b {818}.
    v: Cappelle-Desprez.
Ibf-A1c {818}
    v: Hope.
Ibf-A1d {818}.
    v: Chris.
Ibf-A1e {818}.
    v: Sears' Synthetic.
Ibf-B1 {818}.   5BL {818}. v: CS.
Ibf-B1a {818}.
    v: CS.
Ibf-B1b {818}.
    v: Cappelle-Desprez.
Ibf-B1c {818}.
    v: Ciano 67.
Ibf-B1d {818}.
    v: Sears' Synthetic.
Ibf-D1 {818}.   5DL {818}. v: CS.
Ibf-D1a {818}.
    v: CS.
Ibf-D1b {818}.
    v: Cappelle-Desprez.
Ibf-D1c {818}.
    v: Purple Pericarp.
Ibf-D1d {818}.
    v: Sears' Synthetic.
Ibf-Agi1 {818}.   5Agi {818}.  ad: Vilmorin/Ag. intermedium.
Ibf-E1 {818}.   5EL {818}. ad: CS/Ag. elongatum.
Ibf-H1 {818}.   4H {818}. ad: CS/Betzes.
Ibf-R1 {818}.   5RL {818}. ad: CS/Imperial, CS/KingII.
Ibf-Sl1 {818}.   5Sl {818}. ad: CS/Ae. sharonensis.
Ibf-U1 {818}.   5U {818}. ad: CS/Ae. umbellulata.

5.4. Water soluble proteins

WSP-1 are monomeric grain endosperm proteins identified by their high pI's {817}.
 
Wsp-A1 {817}.   7AL {817}. v: CS.
Wsp-A1a {817}.
    v: CS.
Wsp-A1b {817}.
    v: Huntsman.
Wsp-A1c {817}.
    v: Hope.
Wsp-A1d {817}.
    v: Sicco.
Wsp-A1e {817}.
    v: Condor.
Wsp-B1 {817}.   7BL {817}. v: CS.
Wsp-B1a {817}.
    v: CS.
Wsp-B1b {817}.
    v: Hope.
Wsp-B1c {817}.
    v: Condor.
Wsp-D1 {817}.   7DL {817}. v: CS.
Wsp-D1a {817}.
    v: CS.
Wsp-D1b {817}.
    v: Sears' Synthetic IPS1190903.
Wsp-D1c {893}
    v: T4 = Agatha {893,890}, Indis {890,892}.
Wsp-E1 {817}.   7E {817}. ad: CS/ E. elongata.
Wsp-H1 {817}.   7H {817}. ad: CS/Betzes.
Wsp-Hch1 {817}.   7Hch {817}. ad: CS/H. chilense.
Wsp-Sl1 {817}.   7Sl {817}. ad: CS/Ae. sharonensis.
Wsp-V1 {817}.   7V {817}. ad: CS/D. villosum.

5.5. Salt soluble globulins

GLO-1 are endosperm proteins (23-26 kDa) soluble in salt but not in water {455}.
 
Glo-A1 {455}.   1AS {455}. v: CS.
Glo-B1 {455}.   1BS {455}. v: CS.
Glo-D1 {455}.   1DS {455}. v: CS.

ma: The Glo-1 loci were mapped distally in the short arms of the group 1 chromosomes:
chromosome 1A

Glo-A1 (distal) - 5.2 cM - Gli-A1 {1077}.

Glo-D1 (distal) - 2.9 cM - Gli-D1 {1077}.
 
Glo-E1 {455}.   1ES {455}. ad: CS/E. elongata.
Glo-R1 {455}.   1RS {455}. ad: CS/Imperial.
su: 1B/(1R)E. G., Salzmunde 14/44.

5.6. Waxy proteins

Waxy protein (granule-bound starch synthase = ADP glucose starch glycosyl transferase, EC 2.4 1.21 = GBSS) is tightly bound within endosperm starch granules and is involved in the synthesis of amylase {1616}. Waxy variants, characterised by starch granules containing increased amylopectin and reduced amylose, are preferred for Japaness white salted or "udon" noodles {1650}. Similar waxy phenotypes are controlled by orthologous genes in barley, maize and rice but are not known to occur in rye {725}.
 
Wx-A1 [{180,1053}]. [Xwx-7A {179,180}, Wx-B1 {1053,1054}]. 7AS {180,1053}. v: CS.
Wx-A1a [{1054}].
[Wx-B1a {1054}].   v: CS; Hoshuu.
Wx-A1b [{1054}].
[Wx-B1b {1054}]. Null allele. v: Kanto 79; Kanto 107 Shirodaruma {1617}; Sturdy {1617}.
Wx-A1c {1617}.
    v: QT105 {1617}; WB6 {1617}.
Wx-Ald {1616}.
    tv: T. dicoccoides KU 8937B {1616}.
Wx-Ale {1616}.
    tv: T. durum KU 3655 and KU 3659 {1616}.
Wx-B1 [{180,1053}]. [XWx-4B {179,180}, XWx-4A {961}, Wx-A1 {1053,1054}]. 4AL {180,1054}. v: CS.
Wx-B1a [{1054}].
[Wx-A1a {1054}].   v: CS; Joshuu.
Wx-B1b [{1054}]. 
 [Wx-A1b {1054}]. Null allele. v: Kanto 79; Kanto 82; Kanto 107; Norin 98 Gabo {1617}; Satanta {1617}. For list of Australian wheats, see {1650}.
Wx-B1c {1617}.
    v: Cikataba {1617}; Junbuk 12 {1617}.
Wx-Bld {1616}.
    tv: T. durum KU 4213D and KU 4224C {1616}.
Wx-D1 [{180}], {1053}. [XWx-7D {179,180}]. 7DS {180,1053}. v: CS.
Wx-D1a {1054}.
    v: CS.
Wx-D1b {1617}.
  Null allele. v: Bai Huo {1617}.
Wx-D1c {1617}.
    v: Scoutland {1617}

5.7. Starch granule proteins
 
Sgp-A1 {1615}.   7AS {1615}. v: CS.
Sgp-A1a {1615}.
    v: CS.
Sgp-A1b {1615}.
  Null allele. v: Chosen 30, Chosen 57
Sgp-A1c {1615}.
    v: Hua Non 9.
Sgp-B1 {1615}.   7BS {1615}. v: CS.
Sgp-B1a {1615}.
    v: CS.
Sgp-B1b {1615}.
  Null allele. v: K79.
Sgp-B1c {1615}.
    v: Gnatruche.
Sgp-B1d {1615}.
    v: Waratah.
Sgp-D1 {1615}.   7DS {1615}. v: CS.
Sgp-D1a {1615}.
    v: CS.
Sgp-D1b {1615}.
  Null allele. v: T116.
Sgp-A2 {1615}.     v: CS.
Sgp-B2 {1615}.     v: CS.
Sgp-D2 {1615}.     v: CS.
Sgp-A3 {1615}.   7AS {1615}. v: CS.
Sgp-A3a {1615}.
    v: CS
Sgp-A3b {1615}.
  Null allele. v: Norin 61.
Sgp-B3 {1615}.   7BS {1615}. v: CS.
Sgp-B3a {1615}.
    v: CS.
Sgp-B3b {1615}.
  Null allele. v: Crest.
Sgp-B3c {1615}.
    v: Spica.
Sgp-D3 {1615}.   7DS {1615}. v: CS.

Deletion mapping indicated that the gene order on the 7S arms is; centromere - Sgp-1 - Sgp-3 - Wx{1615}.

5.8. Puroindolines

Puroindolines a and b are the major components of friabilin, a protein complex that is associated with grain texture (see 'Grain Hardness'). Hard wheats result from unique changes in the puroindoline amino acid sequence or (currently) a null form of one of the completely linked genes (max. map distance 4.3 cM) {452}.
 
Pina-D1 {452} 5DS {452}. v: CS
Pina-D1a {452}. v: CS {452}; Heron {1035}.
Pina-D1a is present in all soft hexaploid wheats and possibly all hard hexaploid wheats carrying the Pinb-D1b mutation {452,1035}.
Pina-D1b {1035}. Null allele. v: Falcon {1035}; Butte 86 {1035}.
Pina-D1b may be present in all hard hexaploid wheats not carrying the Pinb-D1b mutation {452,1035}.
Pinb-D1 5DS {452}. v: CS.
Pinb-D1a {452}. v: CS {452}; Hill 81 {452}.
Pinb-D1a is present in all soft hexaploid wheats and possibly all hard hexaploid wheats carrying the Pina-D1b mutation {452,1035}.
Pinb-D1b {452}. 5DS {452} s: CS*/Cheyenne 5D{452}.
v: Wanser {452}.
Pinb-D1b may be present in all hard hexaploid wheats not carrying the Pina-D1b (null) mutation {452,1035}. Wheats with Pinb-D1b contain a Gly-46 to Ser-46 change in amino acid sequence {452}.