Nelson JC et al. (1995) Molecular mapping of wheat. Homoeologous group 3. Genome 38:525-533.
Grain color was scored in the Synthetic x Opata RI population.
Grain color was rated visually, after brief soaking in weak KOH solution, on a scale of 0 to 3 with reference to cultivars Augusta, Karl, and Caldwell, supposed to possess respectively 0, 1, and 2 R genes.