From David Laurie, 7/94
 
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 Reference: DA Laurie, N Pratchett, KM Devos, IJ Leitch & MD Gale
 The distribution of RFLP markers on chromosome 2(2H) of barley in
 relation to the physical and genetic location of 5S rDNA.
 Theor. Appl. Genet. 87: 177-183 (1993)
 
 This paper gives maps of barley chromosome 2 from two crosses.  Eleven
 probes were mapped in both crosses, so the maps can be aligned reasonably
 well.  This paper shows the map in relation to the physical position of
 the 5S genes which was determined using in situ hybridization by Leitch and
 Heslop-Harrison Genome 36: 517-523 (1993).  We also used their nomenclature
 for the sites.
 
 Cross 1: Betzes x Golden Promise
 120 F2 plants
 
 Locus name       cM  distance (Kosambi mapping function)
 Xpsb31a
                 0.4
 Xpsb31b
                 6.0
 Xpsr566
                 3.0
 Xpsr150
                 0
 XMWG878
                 0.4
 XBCD175
                10.3
 Xpsr109
                 1.7
 Xpsr666
                 7.2
 Xpsr108
                 9.6
 Xpsb9
                39.5  (this interval includes the centromere) 
 Xpsr102
                 0
 XCDO366
                 0
 XCDO588
                20.8
 X5SDna-H3
                 3.4
 Xpsr687
                 3.4
 Xpsr901
                 1.5
 Xpsr630
                22.9
 Xpsr540
                 0
 XCDO373
                 6.1
 XBCD266
                30.4
 XWG645
 
 
 Cross 2: Captain x Hordeum spontaneum
 120 F2 plants
 
 Locus name       cM  distance (Kosambi mapping function)
 Xpsb31          
                 3.1
 Xpsr566
                 3.7
 Xpsr150
                12.7
 Xpsr109
                 5.2
 XMWG858
                 3.8
 Xpsb9
                20.0
 Xpsb24
                20.1
 Xpsb30
                 0.6
 XSs2
                 2.1
 Xpsr126
                 5.5 (this interval includes the centromere)
 XWG996
                 9.3
 Xpsr102
                 0
 XCDO588
                18.0
 XMWG865
                 2.7
 X5SDna-H3
                 2.7
 Xpsb141
                10.0
 Xpsr901
                 0.4
 Xpsr630
                 8.8
 Xpsr331
                22.5
 XBCD266
                26.9
 XBCD410
                 3.1
 XWG645
 
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 Reference: DA Laurie, N Pratchett, C Romero, E Simpson & JW Snape
 Assignment of the denso dwarfing gene to the long arm of chromosome 3(3H)
 of barley by use of RFLP markers.
 Plant Breeding 111: 198-203 (1993).
 
 Cross: Magnum x Goldmarker
 113 doubled haploid lines (derived by the Hordeum bulbosum system).
 
 Locus name       cM  distance (Kosambi mapping function)
 XWG940
                 0.0
 Xpsr78
                23.8
 Xpsr74
                 0.0
 Xpsb96
                 4.0
 Xpsr170
                 7.8
 denso
                44.8
 Xpsb177
 
 All these markers are on the long arm.
 
 The denso dwarfing gene has been widely used in European barley breeding.
 It confers a prostrate juvenile growth habit,
 later ear emergence time (heading date)
 and reduced plant height.
 It also gives smaller grains and tends to be associated with lower
 malting quality, though many malting varieties have the gene.
 
 ----------------------------------
 Reference: N Pratchett & DA Laurie
 Genetic map location of the barley developmental mutant liguleless
 in relation to RFLP markers.
 Hereditas 120: 35-39 (1994)
 
 This paper used a bulk segregant analysis to locate the barley 
 li mutation.
 
 Cross : Tayeh-13 (a Japanese variety carrying liguleless) x Betzes
 250 F2 plants from which 36 with the liguleless phenotype were 
 characterized with markers.
 
 Locus name       cM  distance (Kosambi mapping function)
 XCDO366
                  16.4
 hex-v
                  11.4
 Xpsr901
                   3.5
 Xpsr331
                  16.5
 XCDO373
                   4.1
 XBCD266
                   1.3
 liguleless
                  23.9
 XBCD410
                   5.7
 XWG645
 
 
 All these markers are on the long arm.
 
 hex-v controls the 2-row/6-row ear type.  It cosegregates with XMWG865
 - see other group 2 maps (Andreas Graner pers. comm).
 
 ---------------------------------------------------------
 Reference: DA Laurie, N Pratchett, JH Bezant and JW Snape.
 Genetic analysis of a photoperiod response gene on the short arm of
 chromosome 2(2H) of barley (Hordeum vulgare).
 Heredity 72:619-627 (1994)
 
 A photoperiod response gene (designated Ppd-H1) was mapped.  It affects
 flowering time under long days (13h or 16h light) but has little 
 effect under short days (10h).  In the field it is the major determinant
 of flowering time variation in the doubled haploid population.
 Its map location suggests it may be equivalent to the Ppd series of
 photoperiod response genes in wheat.
 This cross also has a second gene (a QTL) affecting flowering time 
 on this chromosome.  However, this gene seems to affect heading data
 pretty well independently of day length.  The second gene is probably
 in the Xpsb24-Xpsr571 interval 
 
 Cross 1: Igri x Triumph
 94 doubled haploid lines (derived by the Hordeum bulbosum method).
 
 Locus name       cM  distance (Kosambi mapping function)
 XMWG878
                  17.7
 Xpsr666
                   2.1
 Xpsr108
                   1.7
 XMWG858
                   1.0
 Ppd-H1
                   5.1
 Xpsb9
                  20.9
 Xpsb24
                  21.1 (the centromere is in this interval)
 Xpsr571
                  31.4
 XMWG865
                  26.5
 Xpsr87
                   0.0
 Xpsr331
                  26.6
 XBCD266
                  28.1
 XBCD410
 
 
 Final note:
 All the map distances are from MAPMAKER.
 
 psr markers are from the Cambridge Lab wheat RFLP group.
 psb markers are our own barley clones.  They are PstI genomic
 fragments cloned in pUC18.  (In the first papers they were called
 psrB markers, but Mike Gale has since persuaded me to change to a
 three letter symbol).
 
 BCD, CDO and WG probes were mapped in barley by Manfred Heun et al.
 (Genome 34: 437-447) and were kindly sent to me by Mark Sorrells.
 MWG probes are from Andreas Graner.  Most are on his map - Graner
 et al. Theor. Appl. Genet. 83: 250-256 (1991).
 
 The common markers allow the maps to be aligned with our wheat and rye
 maps and with the barley maps from Cornell, NABGMP and Germany.